TUESDAY’s CODE

Data: Figure 5 (plus 10 years)

Make data vectors, calculate lambda, and put together dataframe with all necessary data.

census

The census period; an index from 1 to 39 of how many years of data have been collected.

census <- c(1:39)

year t

The year: 1959 to 1997 (Dennis et al use 1959-1987)

year.t <- c(1959:1997)

Population size

Population size is recorded as the number of females with …

females.N <- c(44,47,46,44,46,
               45,46,40,39,39,
               42,39,41,40,33,
               36,34,39,35,34,
               38,36,37,41,39,
               51,47,57,48,60,
               65,74,69,65,57,
               70,81,99,99)

Population growth rate: example

Population growth rate is…

Enter the population size for each year

females.N.1959 <- 44
females.N.1960 <- 47

Calculate the ratio of the 2 population sizes

lambda.59_60 <- females.N.1960/females.N.1959

Access the population sizes by using bracket notation rather than hard coding

# Access the data
females.N[1]
#> [1] 44
females.N[2]
#> [1] 47

# store in objects
females.N.1959 <- females.N[1]
females.N.1960 <- females.N[2]

# confirm the output
females.N.1960/females.N.1959
#> [1] 1.068182

Calculate lambda using bracket notation

lambda.59_60 <- females.N[2]/females.N[1]

The first year of data is 1959. What is lambda for 1958 to 1959?

females.N[1]
#> [1] 44
lambda.58_59 <- females.N[1]/females.N[0]

Population growth rate: vectorized

TASK

Briefly describe (1-2 sentence) what this code is doing.

It calculates two lambdas, of 1959 to 1960 and 1960 to 1961 respectively.

females.N[2:3]
#> [1] 47 46
females.N[1:2]
#> [1] 44 47

females.N[2:3]/females.N[1:2]
#> [1] 1.0681818 0.9787234

This is similar t the previous code chunk, just using all of the data (no need to describe)

length(females.N)
#> [1] 39
females.N[2:39]/females.N[1:38]
#>  [1] 1.0681818 0.9787234 0.9565217 1.0454545 0.9782609 1.0222222 0.8695652
#>  [8] 0.9750000 1.0000000 1.0769231 0.9285714 1.0512821 0.9756098 0.8250000
#> [15] 1.0909091 0.9444444 1.1470588 0.8974359 0.9714286 1.1176471 0.9473684
#> [22] 1.0277778 1.1081081 0.9512195 1.3076923 0.9215686 1.2127660 0.8421053
#> [29] 1.2500000 1.0833333 1.1384615 0.9324324 0.9420290 0.8769231 1.2280702
#> [36] 1.1571429 1.2222222 1.0000000

TASK What does this do? Briefly describe in 1 to 2 sentences why I am using length().

This does the same thing as previous chunk, but is more generalized since we use length to indicate the total number of years.

len <- length(females.N)
females.N[2:len]/females.N[1:len-1]
#>  [1] 1.0681818 0.9787234 0.9565217 1.0454545 0.9782609 1.0222222 0.8695652
#>  [8] 0.9750000 1.0000000 1.0769231 0.9285714 1.0512821 0.9756098 0.8250000
#> [15] 1.0909091 0.9444444 1.1470588 0.8974359 0.9714286 1.1176471 0.9473684
#> [22] 1.0277778 1.1081081 0.9512195 1.3076923 0.9215686 1.2127660 0.8421053
#> [29] 1.2500000 1.0833333 1.1384615 0.9324324 0.9420290 0.8769231 1.2280702
#> [36] 1.1571429 1.2222222 1.0000000

TASK What does this do? Briefly describe in 1 to 2 sentences what is different about this code chunk from the previous one.

This does the same thing as previous code chunk but skips the step of setting and storing length of the population vector to another variable.

females.N[2:length(females.N)]/females.N[1:length(females.N)-1]
#>  [1] 1.0681818 0.9787234 0.9565217 1.0454545 0.9782609 1.0222222 0.8695652
#>  [8] 0.9750000 1.0000000 1.0769231 0.9285714 1.0512821 0.9756098 0.8250000
#> [15] 1.0909091 0.9444444 1.1470588 0.8974359 0.9714286 1.1176471 0.9473684
#> [22] 1.0277778 1.1081081 0.9512195 1.3076923 0.9215686 1.2127660 0.8421053
#> [29] 1.2500000 1.0833333 1.1384615 0.9324324 0.9420290 0.8769231 1.2280702
#> [36] 1.1571429 1.2222222 1.0000000

Negative indexing

Make a short vector to play with; first 10 years

females.N[1:10]
#>  [1] 44 47 46 44 46 45 46 40 39 39
females.Ntemp <- females.N[1:10]

Check - are there 10 numbers

length(females.Ntemp)
#> [1] 10

TASK

What does this do? Briefly describe what the [-1] is doing.

This drops the first element of the list. Negative index drops the element indicated by numeric index. Doesn’t alter the actual content of the vector (i.e. the first element still exists).

females.Ntemp[-1]
#> [1] 47 46 44 46 45 46 40 39 39

TASK How many lambdas can I calculate using the first 10 years of data?

9 lambdas.

females.Ntemp[2:10]/females.Ntemp[1:9]
#> [1] 1.0681818 0.9787234 0.9565217 1.0454545 0.9782609 1.0222222 0.8695652
#> [8] 0.9750000 1.0000000

“Negative indexing” allows you to drop a specific element from a vector.

TASK Drop the the first element

females.Ntemp[-1]
#> [1] 47 46 44 46 45 46 40 39 39

TASK Drop the second element

females.Ntemp[-2]
#> [1] 44 46 44 46 45 46 40 39 39

TASK

How do you drop the 10th element? Type in the code below.

females.Ntemp[-10]
#> [1] 44 47 46 44 46 45 46 40 39

TASK How do you access the last element? Do this in a general way without hard-coding.

females.Ntemp[length(females.Ntemp)]
#> [1] 39

TASK How do DROP the last element? Do this in a general way without hard-coding. By general, I mean in a way that if the length of the vector females.Ntemp changed the code would still drop the correct element.

females.Ntemp[-length(females.Ntemp)]
#> [1] 44 47 46 44 46 45 46 40 39

TASK Calculate the first 9 lambdas.

lambda.i <- females.Ntemp[2:length(females.Ntemp)]/females.Ntemp[1:length(females.Ntemp)-1]

Converting between these 2 code chunks would be a good test question : )

lambda.i <- females.Ntemp[-1]/females.Ntemp[-length(females.Ntemp)]

Calcualte lambdas for all data

TASK

Below each bulleted line describe what the parts of the code do. Run the code to test it.

What does females.N[-1] do? Drops the first element of the vector.

females.N[-1]
#>  [1] 47 46 44 46 45 46 40 39 39 42 39 41 40 33 36 34 39 35 34 38 36 37 41 39 51
#> [26] 47 57 48 60 65 74 69 65 57 70 81 99 99

What does females.N[-length(females.N)] do? Drops the last element of the vector (generalized method using length).

females.N[-length(females.N)]
#>  [1] 44 47 46 44 46 45 46 40 39 39 42 39 41 40 33 36 34 39 35 34 38 36 37 41 39
#> [26] 51 47 57 48 60 65 74 69 65 57 70 81 99

TASK Calculate lambdas for all of the data

females.N[-1]
#>  [1] 47 46 44 46 45 46 40 39 39 42 39 41 40 33 36 34 39 35 34 38 36 37 41 39 51
#> [26] 47 57 48 60 65 74 69 65 57 70 81 99 99
females.N[-length(females.N)]
#>  [1] 44 47 46 44 46 45 46 40 39 39 42 39 41 40 33 36 34 39 35 34 38 36 37 41 39
#> [26] 51 47 57 48 60 65 74 69 65 57 70 81 99

lambda.i <- females.N[-1]/females.N[-length(females.N)]
lambda.i
#>  [1] 1.0681818 0.9787234 0.9565217 1.0454545 0.9782609 1.0222222 0.8695652
#>  [8] 0.9750000 1.0000000 1.0769231 0.9285714 1.0512821 0.9756098 0.8250000
#> [15] 1.0909091 0.9444444 1.1470588 0.8974359 0.9714286 1.1176471 0.9473684
#> [22] 1.0277778 1.1081081 0.9512195 1.3076923 0.9215686 1.2127660 0.8421053
#> [29] 1.2500000 1.0833333 1.1384615 0.9324324 0.9420290 0.8769231 1.2280702
#> [36] 1.1571429 1.2222222 1.0000000

Finish putting together dataframe

Create special columns

TASK

What does this code do? Why do I include NA in the code? (I didn’t cover this in lecture, so just type 1 line - your best guess. “I don’t know” is fine.)

This code seems to append an “empty” value to the end of the lambda.i vector. (I printed out lambda.i to see what it looked like)

lambda.i <- c(lambda.i,NA)

TASK

Check the help file; what type of log does log() calculate (I forgot to put this question on the test!) log() computes logarithm.

lambda_log <- log(lambda.i)

Assemble the dataframe

bear_N <- data.frame(census,
                year.t,
                females.N,
                lambda.i, 
                lambda_log)

TASK

List 3 functions that allow you to examine this dataframe.

summary()
dim()
head()

Examing the population growth rates

Plotting the raw data

TASK

Plot a time series graph of the number of bears (y) versus time (x)
Label the y axis “Population index (females + cubs)”
Label the x axis “Year”
Change the plot to type = “b” so that both points and dots are shown.

plot(females.N~year.t,
     ylab = "Population index (females+cubs)",
     xlab = "Year",
     type = "b")

Bears love to eat trash. Yellowstone closed the last garbage dump in 1970 https://www.yellowstonepark.com/things-to-do/yellowstone-bears-no-longer-get-garbage-treats

THURSDAY’s CODE

census <- 1:39
year.t   <- 1959:1997
females.N <- c(44,47,46,44,46,
               45,46,40,39,39,
               42,39,41,40,33,
               36,34,39,35,34,
               38,36,37,41,39,
               51,47,57,48,60,
               65,74,69,65,57,
               70,81,99,99)
lambda.i <- females.N[-1]/females.N[-length(females.N)]
lambda.i <- c(lambda.i,NA)
lambda_log <- log(lambda.i)
bear_N <- data.frame(census,
                year.t,
                females.N,
                lambda.i, 
                lambda_log)

plot(females.N ~ year.t, data = bear_N, 
     type = "b",
     ylab = "Population index (females + cubs)",
     xlab = "Year")
abline(v = 1970)

How do we determine if a population is likely to go extinct?

We observe the pattern of population growth rate over time. We also model population dynamics by generating random population growth rates and running simulations.

hist(bear_N$lambda.i)

hat_of_lambdas <- bear_N$lambda.i

NA = not applicable, blank, no value

is.na(hat_of_lambdas) # check if something is NA
#>  [1] FALSE FALSE FALSE FALSE FALSE FALSE FALSE FALSE FALSE FALSE FALSE FALSE
#> [13] FALSE FALSE FALSE FALSE FALSE FALSE FALSE FALSE FALSE FALSE FALSE FALSE
#> [25] FALSE FALSE FALSE FALSE FALSE FALSE FALSE FALSE FALSE FALSE FALSE FALSE
#> [37] FALSE FALSE  TRUE

any(is.na(hat_of_lambdas) == TRUE)
#> [1] TRUE

Drop the NA

length(hat_of_lambdas)
#> [1] 39
hat_of_lambdas[39]
#> [1] NA
hat_of_lambdas[-39]
#>  [1] 1.0681818 0.9787234 0.9565217 1.0454545 0.9782609 1.0222222 0.8695652
#>  [8] 0.9750000 1.0000000 1.0769231 0.9285714 1.0512821 0.9756098 0.8250000
#> [15] 1.0909091 0.9444444 1.1470588 0.8974359 0.9714286 1.1176471 0.9473684
#> [22] 1.0277778 1.1081081 0.9512195 1.3076923 0.9215686 1.2127660 0.8421053
#> [29] 1.2500000 1.0833333 1.1384615 0.9324324 0.9420290 0.8769231 1.2280702
#> [36] 1.1571429 1.2222222 1.0000000
hat_of_lambdas[-length(hat_of_lambdas)]
#>  [1] 1.0681818 0.9787234 0.9565217 1.0454545 0.9782609 1.0222222 0.8695652
#>  [8] 0.9750000 1.0000000 1.0769231 0.9285714 1.0512821 0.9756098 0.8250000
#> [15] 1.0909091 0.9444444 1.1470588 0.8974359 0.9714286 1.1176471 0.9473684
#> [22] 1.0277778 1.1081081 0.9512195 1.3076923 0.9215686 1.2127660 0.8421053
#> [29] 1.2500000 1.0833333 1.1384615 0.9324324 0.9420290 0.8769231 1.2280702
#> [36] 1.1571429 1.2222222 1.0000000

na.omit(hat_of_lambdas)
#>  [1] 1.0681818 0.9787234 0.9565217 1.0454545 0.9782609 1.0222222 0.8695652
#>  [8] 0.9750000 1.0000000 1.0769231 0.9285714 1.0512821 0.9756098 0.8250000
#> [15] 1.0909091 0.9444444 1.1470588 0.8974359 0.9714286 1.1176471 0.9473684
#> [22] 1.0277778 1.1081081 0.9512195 1.3076923 0.9215686 1.2127660 0.8421053
#> [29] 1.2500000 1.0833333 1.1384615 0.9324324 0.9420290 0.8769231 1.2280702
#> [36] 1.1571429 1.2222222 1.0000000
#> attr(,"na.action")
#> [1] 39
#> attr(,"class")
#> [1] "omit"

hat_of_lambdas <- hat_of_lambdas[-length(hat_of_lambdas)]

hist(hat_of_lambdas)

Random sampling of lambda

Draws a sample of values of lambda from the pool/hat with replacement. (Therefore repeats are possible)

# randomly choosing a lamda value from the hat
sample(x = hat_of_lambdas, 
       size = 1,
       replace = TRUE)  # reuses data from the same hat
#> [1] 1.222222

# saving the pulled lamda to variable
lambda_rand.t <- sample(x = hat_of_lambdas, size = 1, replace = TRUE)
lambda_rand.t
#> [1] 1

Getting initial population size

Shows different information about the data set. The simulation starts at the year 1997.

head(bear_N)
#>   census year.t females.N  lambda.i  lambda_log
#> 1      1   1959        44 1.0681818  0.06595797
#> 2      2   1960        47 0.9787234 -0.02150621
#> 3      3   1961        46 0.9565217 -0.04445176
#> 4      4   1962        44 1.0454545  0.04445176
#> 5      5   1963        46 0.9782609 -0.02197891
#> 6      6   1964        45 1.0222222  0.02197891
tail(bear_N)
#>    census year.t females.N  lambda.i lambda_log
#> 34     34   1992        65 0.8769231 -0.1313360
#> 35     35   1993        57 1.2280702  0.2054440
#> 36     36   1994        70 1.1571429  0.1459539
#> 37     37   1995        81 1.2222222  0.2006707
#> 38     38   1996        99 1.0000000  0.0000000
#> 39     39   1997        99        NA         NA
summary(bear_N)
#>      census         year.t       females.N        lambda.i     
#>  Min.   : 1.0   Min.   :1959   Min.   :33.00   Min.   :0.8250  
#>  1st Qu.:10.5   1st Qu.:1968   1st Qu.:39.00   1st Qu.:0.9452  
#>  Median :20.0   Median :1978   Median :44.00   Median :1.0000  
#>  Mean   :20.0   Mean   :1978   Mean   :49.79   Mean   :1.0281  
#>  3rd Qu.:29.5   3rd Qu.:1988   3rd Qu.:57.00   3rd Qu.:1.1038  
#>  Max.   :39.0   Max.   :1997   Max.   :99.00   Max.   :1.3077  
#>                                                NA's   :1       
#>    lambda_log      
#>  Min.   :-0.19237  
#>  1st Qu.:-0.05639  
#>  Median : 0.00000  
#>  Mean   : 0.02134  
#>  3rd Qu.: 0.09874  
#>  Max.   : 0.26826  
#>  NA's   :1
dim(bear_N)
#> [1] 39  5
N.1997 <- 99

One round of population simulation

Calculate the number of bears in 1998 and set it as a variable.

1.22807*99
#> [1] 121.5789
lambda_rand.t*N.1997
#> [1] 99
N.1998 <- lambda_rand.t*N.1997

Simulation the hard way

Not very good way to simulate.

# 1997-1998
lambda_rand.t <- sample(x = hat_of_lambdas, size = 1,replace = TRUE)
N.1998 <- lambda_rand.t*N.1997

# 1998-1999
lambda_rand.t <- sample(x = hat_of_lambdas, size = 1,replace = TRUE)
N.1999 <- lambda_rand.t*N.1998

# 1999-2000
lambda_rand.t <- sample(x = hat_of_lambdas, size = 1,replace = TRUE)
N.2000 <- lambda_rand.t*N.1999

# 2000-2001
lambda_rand.t <- sample(x = hat_of_lambdas, size = 1,replace = TRUE)
N.2001 <- lambda_rand.t*N.2000

# 2001-2002
lambda_rand.t <- sample(x = hat_of_lambdas, size = 1,replace = TRUE)
N.2002 <- lambda_rand.t*N.2001

# 2002-2003
lambda_rand.t <- sample(x = hat_of_lambdas, size = 1,replace = TRUE)
N.2003 <- lambda_rand.t*N.2002

# 2003-2004
lambda_rand.t <- sample(x = hat_of_lambdas, size = 1,replace = TRUE)
N.2004 <- lambda_rand.t*N.2003

# 2004-2005
lambda_rand.t <- sample(x = hat_of_lambdas, size = 1,replace = TRUE)
N.2005 <- lambda_rand.t*N.2004

Plotting the simulation result

Using two vectors, year and generated numbers of bears to make a data frame. Plot a scatter plot.

year <- seq(1997, 2004)
N.rand <- c(N.1998,N.1999,N.2000,N.2001,N.2002,N.2003,N.2004,N.2005)
df.rand <- data.frame(N.rand, year)
plot(N.rand ~ year, data = df.rand, type = "b")

Using for loop in simulation

The not using for() loop version

# Initial conditions

N.1997 <- 99
N.initial <- 99

# Explore xlim = argument
plot(N.1997 ~ c(1997))

plot(N.1997 ~ c(1997),
     xlim = c(1997, 1997+50))


# xlim and ylim
plot(N.1997 ~ c(1997),
     xlim = c(1997, 1997+50),
     ylim = c(0, 550))

# for loop the hard way (not using for())
#
N.current <- N.initial

# this is where the for() loop would be
t <- 1
  
  # take a lambda
  lambda_rand.t <- sample(x = hat_of_lambdas, size = 1,replace = TRUE)
  
  # determine population size
  N.t <- N.current*lambda_rand.t
  
  # what year we start
  year.t <- 1997+t
  
  # plot population
  # points() updates
  points(N.t ~ year.t)

  
  # update to current
  N.current <- N.t

The using for() loop version.


# makes a new plot with starting population size
plot(N.1997 ~ c(1997), xlim = c(1997, 1997+50), ylim = c(0, 550))

# start at 1997s population size
N.current <- N.1997

# the for() loop
for(t in 1:50){
  
  # at start of new iteration, grab new lambda
  lambda_rand.t <- sample(x = hat_of_lambdas, 
                          size = 1,
                          replace = TRUE)
  
  # determine population size
  N.t <- N.current*lambda_rand.t
  
  # update year
  year.t <- 1997+t
  
  # update points
  points(N.t ~ year.t)
  
  # update population size
  N.current <- N.t
}

R plotting code (magic!) Multiple plots side by side

par(mfrow = c(3,3), # how many rows and columns
    mar = c(1,1,1,1)) # margin size

The for() loop but with restrictions/limits. Use xlim and ylim to set limits on year and carrying capacity.

plot(N.1997 ~ c(1997), xlim = c(1997, 1997+50), ylim = c(0, 550))
N.current <- N.1997
for(t in 1:50){
  
  lambda_rand.t <- sample(x = hat_of_lambdas, 
                          size = 1,
                          replace = TRUE)
  
  N.t <- N.current*lambda_rand.t
  
  year.t <- 1997+t
  
  points(N.t ~ year.t)
  
  N.current <- N.t
}

Plotting Multiple Simulations

The xaxt and yaxt here are set to n which makes no axis labels shown

par(mfrow = c(10,10), mar = c(0,0,0,0), xaxt = "n", yaxt = "n")

Added a red line to keep track. 10x10 simulation. But still not efficient enough.


plot(N.1997 ~ c(1997), xlim = c(1997, 1997+50), ylim = c(0, 550), cex = 0.5)
abline(h = N.1997)
abline(h = 0, col = "red")
N.current <- N.1997
for(t in 1:50){
  lambda_rand.t <- sample(x = hat_of_lambdas, size = 1,replace = TRUE)
  N.t <- N.current*lambda_rand.t
  year.t <- 1997+t
  points(N.t ~ year.t, cex = 0.5)
  
  N.current <- N.t
}

Bear Population Simulation

Di Zhang

9/22/2020

TUESDAY’s CODE

Data: Figure 5 (plus 10 years)

census

year t

Population size

Population growth rate: example

Population growth rate: vectorized

Negative indexing

Calcualte lambdas for all data

Finish putting together dataframe

Create special columns

Assemble the dataframe

Examing the population growth rates

Plotting the raw data

THURSDAY’s CODE

How do we determine if a population is likely to go extinct?

Random sampling of lambda

Getting initial population size

One round of population simulation

Simulation the hard way

Plotting the simulation result

Using for loop in simulation

Plotting Multiple Simulations