Metadata for growth/scales:

Spacing and circuli

FMC (num) circuli number identifying First marine circulus

FMC.spacing (num) FMC intercirculi spacing

marine.growth (num) marine growth = FMC to end

total.growth (num) entire scale from 1st circuli to the end

smolt.circ (num) number of smolt circuli

marine.circ (num) number of marine circuli

total.circ (num) number of total circuli

FSM.spacing (num) first summer maximum intercirculi spacing

M1.spacing (num) M1 intercirculi spacing

FS.circ (num) First summer, number of circuli

FW.circ (num) First winter, number of circuli

PS.circ (num) Post-smolt, number of circuli

SSM.spacing (num) Second summer maximum intercirculi spacing

M2.spacing (num) M2 intercirculi spacing

SS.circ (num) Second summer, number of circuli

SW.circ (num) Second winter, number of circuli

Y2.circ (num) Year-two number of circuli

Increments

smolt.incr (num) smolt increment (focus to last circuli before FMC)

FS.incr (num) first summer at sea growth (FMC to FSM)

FW.incr (num) first winter at sea growth (first circuli after FSM to M1)

PS.incr (num) post-smolt growth (FMC to M1)

SS.incr (num) Second summer at sea growth (first circuli after M1 to SSM (second summer max))

SW.incr (num) Second winter at sea growth (first circuli after SSM to M2)

Y2.incr (num) Year 2 growth (first circuli after M1 to M2)

Carlin tag EDA

Sample sizes

Summary statistics for each incremental growth period based on algorithm calculations.

Increment n Mean SD Min Max
smolt.incr 1661 1.147 0.286 0.299 2.342
FS.incr 1621 0.676 0.250 0.109 2.262
FW.incr 1621 0.845 0.293 0.068 2.590
PS.incr 1621 1.521 0.308 0.592 3.284
SS.incr 135 0.560 0.254 0.049 1.366
SW.incr 135 0.866 0.315 0.095 1.984
Y2.incr 135 1.422 0.326 0.094 2.279

Annual sample size, mean, and sd by sea age for year one increments.

Smolt Increment
First Summer Incr.
First Winter Incr.
Postsmolt Incr.
Release Year Sea Age n Mean SD Mean SD Mean SD Mean SD
1966 1 21 0.96 0.19 0.65 0.31 1.09 0.29 1.74 0.24
1966 2 23 0.85 0.20 0.61 0.35 1.11 0.35 1.72 0.24
1967 0 5 1.03 0.16
1967 1 4 0.95 0.41 0.83 0.23 1.06 0.59 1.89 0.57
1967 2 7 0.93 0.32 0.78 0.36 0.93 0.39 1.71 0.23
1968 0 5 0.96 0.26
1968 1 7 1.10 0.32 0.62 0.28 1.06 0.22 1.68 0.28
1969 1 67 1.10 0.33 0.68 0.26 0.79 0.26 1.47 0.35
1969 2 16 1.08 0.35 0.75 0.33 0.68 0.28 1.43 0.31
1970 1 257 1.22 0.34 0.67 0.23 0.84 0.32 1.51 0.36
1970 2 8 1.12 0.40 0.82 0.32 0.86 0.47 1.67 0.57
1971 1 60 1.26 0.26 0.65 0.2 0.84 0.25 1.5 0.31
1971 2 12 1.27 0.27 0.53 0.16 0.89 0.39 1.42 0.34
1972 0 17 1.29 0.30
1972 1 88 1.20 0.30 0.72 0.27 0.88 0.33 1.6 0.3
1972 2 11 0.83 0.25 0.82 0.18 0.94 0.35 1.75 0.23
1973 1 185 1.24 0.24 0.65 0.26 0.85 0.28 1.5 0.26
1973 2 8 1.37 0.21 0.59 0.13 0.8 0.2 1.39 0.14
1974 0 5 1.31 0.37
1974 1 80 1.19 0.22 0.66 0.23 0.94 0.24 1.6 0.28
1974 2 4 0.93 0.21 0.67 0.32 0.84 0.13 1.51 0.19
1975 1 62 1.11 0.26 0.74 0.26 0.82 0.36 1.56 0.31
1976 1 15 0.97 0.32 0.63 0.28 0.8 0.26 1.43 0.32
1977 1 6 1.36 0.30 0.55 0.15 1 0.32 1.55 0.26
1978 2 5 1.18 0.18 0.66 0.17 0.84 0.23 1.49 0.13
1979 1 159 1.21 0.28 0.72 0.23 0.77 0.22 1.48 0.27
1979 2 7 1.15 0.35 0.82 0.26 0.9 0.17 1.72 0.35
1980 1 24 1.12 0.21 0.54 0.23 0.91 0.35 1.44 0.29
1980 2 8 0.94 0.37 0.9 0.19 0.7 0.13 1.6 0.23
1981 1 34 1.19 0.29 0.68 0.31 0.77 0.28 1.45 0.35
1982 1 15 1.03 0.31 0.62 0.27 0.73 0.27 1.35 0.24
1983 1 28 1.14 0.25 0.66 0.23 0.66 0.17 1.32 0.28
1984 1 69 1.06 0.21 0.71 0.24 0.71 0.23 1.42 0.22
1985 1 62 1.06 0.20 0.59 0.22 0.91 0.26 1.5 0.25
1986 1 95 1.06 0.18 0.72 0.26 0.86 0.25 1.58 0.27
1986 2 7 0.96 0.17 0.67 0.28 0.78 0.14 1.45 0.23
1987 1 48 0.99 0.21 0.64 0.2 0.88 0.32 1.52 0.27
1988 1 68 1.07 0.21 0.71 0.27 0.85 0.27 1.56 0.3
1989 1 6 1.06 0.22 0.78 0.14 1.01 0.16 1.79 0.27
1990 1 11 1.04 0.27 0.58 0.16 0.95 0.27 1.53 0.24

Annual sample size, mean, and sd for second year increments.

Second Summer Incr.
Second Winter Incr.
Year 2 Increment
Release Year Sea Age n Mean SD Mean SD Mean SD
1966 2 19 0.70 0.21 0.73 0.26 1.43 0.29
1967 2 5 0.42 0.06 0.88 0.14 1.30 0.13
1968 3 1 0.78 0.59 1.37
1969 2 15 0.43 0.19 0.90 0.19 1.33 0.25
1970 2 6 0.53 0.31 0.86 0.18 1.39 0.24
1971 2 11 0.66 0.3 0.90 0.32 1.56 0.28
1972 2 11 0.44 0.17 0.93 0.26 1.37 0.23
1973 2 8 0.65 0.28 0.97 0.32 1.62 0.36
1974 2 4 0.49 0.14 1.39 0.4 1.88 0.3
1975 2 1 0.43 1.29 1.71
1976 2 3 0.58 0.07 0.91 0.71 1.46 0.71
1977 2 2 0.48 0.23 0.79 0.2 1.27 0.02
1978 2 5 0.49 0.21 0.81 0.5 1.27 0.4
1979 2 7 0.50 0.2 0.99 0.2 1.49 0.28
1980 2 8 0.63 0.28 0.90 0.38 1.53 0.28
1981 2 1 0.49 1.29 1.78
1982 2 1 0.95 0.75 1.70
1983 2 2 0.30 0.02 0.74 0.06 1.04 0.04
1984 2 1 0.81 0.48 1.29
1986 2 7 0.46 0.09 0.85 0.13 1.31 0.17

As expected, sample size is not evenly distributed among years of the study, but there are peaks in the early 70s and late 80s. Overall, the growth data for 2SW fish and beyond is very limited. Only 215 2SW fish were caught at sea. About 3500 2SW fish returned to the rivers, but we don’t have scale data from “river return” fish (only distant).

Temporal Differences

Annual mean and standard deviations for all growth increments. Blue represents 1SW fish, green represents 2SW fish, which is why there are only green lines for second year increments. I made boxplots as well, but this looked a little cleaner, and I think it shows the same trends.

Time series with 5-year smoothing for first summer growth and post-smolt growth by age at recapture for distant recaptures only.

It was interesting that weight at capture and length at capture was not correlated with the growth increments. The other intersting thing in this plot was that there is a negative relationship between FS and FW increment. Perhaps there is a trade off between growing quickly at first and expending all of your resources versus growing slow and steady all summer and winter?

Spatial differences

I was curious if there were any differences in first winter growth by recapture location. Perhaps some areas are more advantageous for wintering fish. At the state level, it was difficult to see any spatial or temporal differences.

Basic PCA to look differences in first year growth by recapture location.