## Loading required package: readxl
## Loading required package: lubridate
## 
## Attaching package: 'lubridate'
## The following objects are masked from 'package:data.table':
## 
##     hour, isoweek, mday, minute, month, quarter, second, wday, week,
##     yday, year
## The following objects are masked from 'package:base':
## 
##     date, intersect, setdiff, union
## Loading required package: solartime
## - Winter data excluded.
## - The counts of all passing / wintering species (not resident nor breeding in the sampled unit) were set to zero. Abreed_and_non_breed contains the non-breeders as well.
## - Species less likely to be interacting with sampling location were EXCLUDED

Mediterranean Maquis

Unit is divided into 3 subunits: Judea, Carmel and Galilee. Factors are proximity to settlements and time. Sampling started in spring 2012 (pilot year), but during T0 sampling was performed only in winter of 2014, and the next spring sampling was done in T1 (2015). Therefore 2012 will not be considered as pilot here but as T0. Total 5 campaigns, 3subunits per campaign, 5 sites per subunit, with 6 plots per site (total of 450 plot-campaign combinations).

Raw data Total abundance: 21855 Number of observations: 5795 Total richness: 108

Filtered data Total abundance: 13796 Number of observations: 4265 Total richness: 49

Model gma, abundance and richness

Richness done with rare species. Abundance and mean abundance done without rare species. Full models include cosine and sine of the time difference from June 21st (in radians).

richness

Explore data and plot mean-variance plot. There is a strong relationship, indicating that employing GLMs is the proper way to analyze, rather than OLS (assumption of homogeneity is violated).

## [1] "RICHNESS WITH RARE SPECIES"
## ℹ SHA-1 hash of file is "eef55df1963ba201f57cf44513f566298756fd4f"
richness year_ct site settlements subunit td_sc cos_td_rad sin_td_rad h_from_sunrise cos_hsun sin_hsun monitors_name wind precipitation temperature clouds
Min. : 0.000 Min. :0.0 Nir Etzion : 31 Far :224 Judean Highlands:149 Min. :-1.9882 Min. :-0.1671 Min. :-1.0000 Length:444 Min. :-0.3289 Min. :-0.1253 Sassi Haham : 53 0 : 33 0 :119 0 : 4 0 : 32
1st Qu.: 6.000 1st Qu.:3.0 Ein Yaakov : 30 Near:220 Carmel :150 1st Qu.:-0.2713 1st Qu.: 0.5702 1st Qu.:-0.8215 Class :difftime 1st Qu.: 0.6855 1st Qu.: 0.2563 Eyal Shochat: 47 1 : 42 3 : 2 1 : 38 1 : 0
Median : 8.000 Median :5.0 Givat Yearim : 30 NA Galilee :145 Median : 0.4918 Median : 0.8140 Median :-0.5808 Mode :numeric Median : 0.8738 Median : 0.4863 Eran Banker : 30 2 : 12 NA’s:323 2 : 37 2 : 5
Mean : 8.054 Mean :4.8 Givat Yeshayahu: 30 NA NA Mean : 0.3205 Mean : 0.7098 Mean :-0.5864 NA Mean : 0.7997 Mean : 0.4836 Asaf Mayrose: 18 3 : 2 NA 3 : 18 3 : 13
3rd Qu.:10.000 3rd Qu.:7.0 Goren : 30 NA NA 3rd Qu.: 1.1023 3rd Qu.: 0.9413 3rd Qu.:-0.3375 NA 3rd Qu.: 0.9666 3rd Qu.: 0.7281 Ohad Sharir : 18 NA’s:355 NA NA’s:347 NA’s:394
Max. :18.000 Max. :9.0 Kerem Maharal : 30 NA NA Max. : 1.7127 Max. : 0.9976 Max. :-0.0688 NA Max. : 1.0000 Max. : 0.9998 (Other) : 58 NA NA NA NA
NA NA (Other) :263 NA NA NA NA NA NA NA’s :89 NA’s :89 NA’s :220 NA NA NA NA

no observation for all 4 weather variables. many NAs for sampling time of day variables.exclude from model. An extreme observation of richness>20 in Judean highlands near settlements.

richness year_ct site settlements subunit td_sc cos_td_rad sin_td_rad h_from_sunrise cos_hsun sin_hsun
richness 1.0000000 -0.0647635 0.0926545 0.4762118 -0.1936858 0.0617968 0.0751315 0.0460403 -0.0582024 0.0933868 -0.0342460
year_ct -0.0647635 1.0000000 -0.0460962 -0.0041782 0.0046005 -0.6240340 -0.5599811 -0.6642968 0.0244310 -0.0070326 0.0341395
site 0.0926545 -0.0460962 1.0000000 0.0038640 -0.1472070 0.0103683 0.0017250 0.0296647 -0.0105111 -0.0092740 -0.0240278
settlements 0.4762118 -0.0041782 0.0038640 1.0000000 -0.0167082 0.0029695 0.0016285 0.0037459 0.0341209 -0.0071595 0.0507160
subunit -0.1936858 0.0046005 -0.1472070 -0.0167082 1.0000000 -0.0056370 -0.0129685 -0.0136922 0.1193296 -0.1166460 0.1174210
td_sc 0.0617968 -0.6240340 0.0103683 0.0029695 -0.0056370 1.0000000 0.9788766 0.9693667 -0.1975082 0.1811635 -0.2043768
cos_td_rad 0.0751315 -0.5599811 0.0017250 0.0016285 -0.0129685 0.9788766 1.0000000 0.9008724 -0.1878186 0.1700185 -0.1970275
sin_td_rad 0.0460403 -0.6642968 0.0296647 0.0037459 -0.0136922 0.9693667 0.9008724 1.0000000 -0.2005880 0.1868900 -0.2040813
h_from_sunrise -0.0582024 0.0244310 -0.0105111 0.0341209 0.1193296 -0.1975082 -0.1878186 -0.2005880 1.0000000 -0.9598144 0.9818950
cos_hsun 0.0933868 -0.0070326 -0.0092740 -0.0071595 -0.1166460 0.1811635 0.1700185 0.1868900 -0.9598144 1.0000000 -0.8930078
sin_hsun -0.0342460 0.0341395 -0.0240278 0.0507160 0.1174210 -0.2043768 -0.1970275 -0.2040813 0.9818950 -0.8930078 1.0000000

Fit Poisson glm, check for existence of overdispersion

## [1] "Estimating overdispersion parameter phi: (Res. Dev.)/(n-p) where n=number of observations; p=number of parameters in the model."
## [1] "od = 0.917251305665046"

Overdispersion parameter is < 1. Choose Poisson.

##                         GVIF Df GVIF^(1/(2*Df))
## settlements         3.275497  1        1.809834
## year_ct             3.277089  1        1.810273
## subunit             1.001389  2        1.000347
## cos_td_rad          5.439321  1        2.332235
## sin_td_rad          6.692207  1        2.586930
## settlements:year_ct 4.690440  1        2.165742

go with mixed model, attempt model selection yet.

## Generalized linear mixed model fit by maximum likelihood (Laplace
##   Approximation) [glmerMod]
##  Family: poisson  ( log )
## Formula: 
## richness ~ settlements * year_ct + subunit + cos_td_rad + sin_td_rad +  
##     (1 | site)
##    Data: P.anal
## 
##      AIC      BIC   logLik deviance df.resid 
##   2115.0   2151.8  -1048.5   2097.0      435 
## 
## Scaled residuals: 
##      Min       1Q   Median       3Q      Max 
## -3.04280 -0.60906 -0.07994  0.57865  2.87628 
## 
## Random effects:
##  Groups Name        Variance Std.Dev.
##  site   (Intercept) 0.009067 0.09522 
## Number of obs: 444, groups:  site, 16
## 
## Fixed effects:
##                          Estimate Std. Error z value Pr(>|z|)    
## (Intercept)              1.575932   0.201233   7.831 4.83e-15 ***
## settlementsNear          0.393867   0.061554   6.399 1.57e-10 ***
## year_ct                 -0.004072   0.009709  -0.419  0.67493    
## subunitCarmel            0.082972   0.071979   1.153  0.24902    
## subunitGalilee          -0.198418   0.072660  -2.731  0.00632 ** 
## cos_td_rad               0.299159   0.148180   2.019  0.04350 *  
## sin_td_rad              -0.228252   0.169323  -1.348  0.17765    
## settlementsNear:year_ct -0.002007   0.010854  -0.185  0.85329    
## ---
## Signif. codes:  0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1
## 
## Correlation of Fixed Effects:
##             (Intr) sttlmN yer_ct sbntCr sbntGl cs_td_ sn_td_
## settlmntsNr -0.190                                          
## year_ct     -0.027  0.552                                   
## subunitCrml -0.195 -0.002  0.005                            
## subunitGall -0.199  0.003 -0.001  0.501                     
## cos_td_rad  -0.931  0.008 -0.080  0.019  0.019              
## sin_td_rad   0.866 -0.010  0.328 -0.006 -0.018 -0.853       
## sttlmntsN:_  0.158 -0.833 -0.660  0.003  0.003 -0.006  0.009

perform stepwise model selection of poisson mixed model.

## Warning in checkConv(attr(opt, "derivs"), opt$par, ctrl = control$checkConv, :
## Model failed to converge with max|grad| = 0.0035507 (tol = 0.002, component 1)
## Single term deletions
## 
## Model:
## richness ~ settlements * year_ct + subunit + cos_td_rad + sin_td_rad + 
##     (1 | site)
##                     npar    AIC
## <none>                   2115.0
## subunit                2 2122.2
## cos_td_rad             1 2117.1
## sin_td_rad             1 2114.8
## settlements:year_ct    1 2113.0

remove settlements X year.

## Single term deletions
## 
## Model:
## richness ~ settlements + year_ct + subunit + cos_td_rad + sin_td_rad + 
##     (1 | site)
##             npar    AIC
## <none>           2113.0
## settlements    1 2240.8
## year_ct        1 2111.5
## subunit        2 2120.3
## cos_td_rad     1 2115.1
## sin_td_rad     1 2112.8

drop year

## Single term deletions
## 
## Model:
## richness ~ settlements + subunit + cos_td_rad + sin_td_rad + 
##     (1 | site)
##             npar    AIC
## <none>           2111.5
## settlements    1 2239.4
## subunit        2 2118.7
## cos_td_rad     1 2113.3
## sin_td_rad     1 2110.8

drop sine.

## Single term deletions
## 
## Model:
## richness ~ settlements + subunit + cos_td_rad + (1 | site)
##             npar    AIC
## <none>           2110.8
## settlements    1 2238.5
## subunit        2 2118.0
## cos_td_rad     1 2113.5

Settlements and cosin time of year remain. Final model:

## Generalized linear mixed model fit by maximum likelihood (Laplace
##   Approximation) [glmerMod]
##  Family: poisson  ( log )
## Formula: richness ~ settlements + subunit + cos_td_rad + (1 | site)
##    Data: P.anal
## 
##      AIC      BIC   logLik deviance df.resid 
##   2110.8   2135.4  -1049.4   2098.8      438 
## 
## Scaled residuals: 
##      Min       1Q   Median       3Q      Max 
## -3.09026 -0.59675 -0.07933  0.56203  2.96581 
## 
## Random effects:
##  Groups Name        Variance Std.Dev.
##  site   (Intercept) 0.009619 0.09808 
## Number of obs: 444, groups:  site, 16
## 
## Fixed effects:
##                 Estimate Std. Error z value Pr(>|z|)    
## (Intercept)      1.80687    0.07258  24.896   <2e-16 ***
## settlementsNear  0.38421    0.03401  11.296   <2e-16 ***
## subunitCarmel    0.08257    0.07349   1.124   0.2612    
## subunitGalilee  -0.20004    0.07405  -2.702   0.0069 ** 
## cos_td_rad       0.13535    0.06305   2.147   0.0318 *  
## ---
## Signif. codes:  0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1
## 
## Correlation of Fixed Effects:
##             (Intr) sttlmN sbntCr sbntGl
## settlmntsNr -0.279                     
## subunitCrml -0.526  0.002              
## subunitGall -0.508  0.009  0.502       
## cos_td_rad  -0.635 -0.002  0.021 -0.003
## $site
##                   (Intercept)
## Abirim          -7.321848e-02
## Aderet           6.137108e-02
## Beit Oren       -2.233669e-02
## Ein Yaakov      -2.970003e-02
## Givat Yearim    -7.623271e-02
## Givat Yeshayahu  9.364091e-02
## Goren           -7.232404e-03
## Iftach           1.773011e-01
## Kerem Maharal    9.669359e-02
## Kfar Shamai     -5.892993e-02
## Margaliot       -1.725282e-03
## Nehusha         -4.299542e-02
## Nir Etzion      -1.521589e-01
## Ofer             8.260917e-02
## Ramat Raziel    -3.076398e-02
## Yagur            2.684868e-05
## 
## with conditional variances for "site"
## $site

attempt to add year, see if significant:

## Generalized linear mixed model fit by maximum likelihood (Laplace
##   Approximation) [glmerMod]
##  Family: poisson  ( log )
## Formula: richness ~ year_ct + settlements + subunit + cos_td_rad + (1 |  
##     site)
##    Data: P.anal
## 
##      AIC      BIC   logLik deviance df.resid 
##   2112.8   2141.5  -1049.4   2098.8      437 
## 
## Scaled residuals: 
##     Min      1Q  Median      3Q     Max 
## -3.0946 -0.6000 -0.0782  0.5691  2.9651 
## 
## Random effects:
##  Groups Name        Variance Std.Dev.
##  site   (Intercept) 0.009581 0.09788 
## Number of obs: 444, groups:  site, 16
## 
## Fixed effects:
##                   Estimate Std. Error z value Pr(>|z|)    
## (Intercept)      1.8147139  0.0962891  18.847   <2e-16 ***
## year_ct         -0.0008103  0.0065499  -0.124   0.9015    
## settlementsNear  0.3841894  0.0340127  11.295   <2e-16 ***
## subunitCarmel    0.0824132  0.0733925   1.123   0.2615    
## subunitGalilee  -0.2001589  0.0739537  -2.707   0.0068 ** 
## cos_td_rad       0.1298876  0.0769263   1.688   0.0913 .  
## ---
## Signif. codes:  0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1
## 
## Correlation of Fixed Effects:
##             (Intr) yer_ct sttlmN sbntCr sbntGl
## year_ct     -0.658                            
## settlmntsNr -0.214  0.005                     
## subunitCrml -0.407  0.016  0.002              
## subunitGall -0.391  0.013  0.009  0.502       
## cos_td_rad  -0.769  0.574  0.001  0.027  0.005

year not significant, rightfully dropped. center time of year variable, highly correlated with intercept.

## Generalized linear mixed model fit by maximum likelihood (Laplace
##   Approximation) [glmerMod]
##  Family: poisson  ( log )
## Formula: richness ~ settlements + subunit + cos_td_rad_c + (1 | site)
##    Data: P.anal
## 
##      AIC      BIC   logLik deviance df.resid 
##   2110.8   2135.4  -1049.4   2098.8      438 
## 
## Scaled residuals: 
##      Min       1Q   Median       3Q      Max 
## -3.09026 -0.59674 -0.07933  0.56201  2.96580 
## 
## Random effects:
##  Groups Name        Variance Std.Dev.
##  site   (Intercept) 0.009619 0.09808 
## Number of obs: 444, groups:  site, 16
## 
## Fixed effects:
##                 Estimate Std. Error z value Pr(>|z|)    
## (Intercept)      1.90293    0.05611  33.916   <2e-16 ***
## settlementsNear  0.38421    0.03401  11.296   <2e-16 ***
## subunitCarmel    0.08256    0.07349   1.124   0.2612    
## subunitGalilee  -0.20004    0.07404  -2.702   0.0069 ** 
## cos_td_rad_c     0.13534    0.06305   2.147   0.0318 *  
## ---
## Signif. codes:  0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1
## 
## Correlation of Fixed Effects:
##             (Intr) sttlmN sbntCr sbntGl
## settlmntsNr -0.363                     
## subunitCrml -0.663  0.002              
## subunitGall -0.660  0.009  0.502       
## cos_td_rd_c -0.023 -0.002  0.021 -0.003
## Registered S3 methods overwritten by 'broom':
##   method            from  
##   tidy.glht         jtools
##   tidy.summary.glht jtools
Observations 444
Dependent variable richness
Type Mixed effects generalized linear model
Family poisson
Link log
AIC 2110.815
BIC 2135.390
Pseudo-R² (fixed effects) 0.292
Pseudo-R² (total) 0.345
Fixed Effects
exp(Est.) S.E. z val. p
(Intercept) 6.706 0.056 33.916 0.000
settlementsNear 1.468 0.034 11.296 0.000
subunitCarmel 1.086 0.073 1.124 0.261
subunitGalilee 0.819 0.074 -2.702 0.007
cos_td_rad_c 1.145 0.063 2.147 0.032
Random Effects
Group Parameter Std. Dev.
site (Intercept) 0.098
Grouping Variables
Group # groups ICC
site 16 0.010
## Loading required package: Cairo
## Loading required package: extrafont
## Registering fonts with R
## Confidence intervals for merMod models is an experimental feature. The
## intervals reflect only the variance of the fixed effects, not the random
## effects.

##        2 
## 3.141228
##        2 
## 46.84533
##        1        2 
## 6.705529 9.846757
## [1] 0.4684534
## Confidence intervals for merMod models is an experimental feature. The
## intervals reflect only the variance of the fixed effects, not the random
## effects.

##  subunit          emmean     SE  df asymp.LCL asymp.UCL
##  Judean Highlands   2.10 0.0524 Inf      1.99      2.20
##  Carmel             2.18 0.0518 Inf      2.08      2.28
##  Galilee            1.89 0.0526 Inf      1.79      2.00
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## Confidence level used: 0.95
## [1] 33.64275
## [1] 8.328707
## [1] 23.36781
##  contrast                   estimate     SE  df z.ratio p.value
##  Judean Highlands - Carmel   -0.0826 0.0735 Inf  -1.124  0.2612
##  Judean Highlands - Galilee   0.2000 0.0740 Inf   2.702  0.0103
##  Carmel - Galilee             0.2826 0.0736 Inf   3.839  0.0004
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## P value adjustment: fdr method for 3 tests

statistically significant lower richness in Galilee and far from settlements. No significant change in richness over time.

geometric mean of abundance

Explore data. Exclude time of day because of high number of NAs.

## [1] "GEOMETRIC MEAN ABUNDANCE WITHOUT RARE SPECIES"
## Warning: Removed 1 rows containing non-finite values (`stat_boxplot()`).
## Removed 1 rows containing non-finite values (`stat_boxplot()`).
## Removed 1 rows containing non-finite values (`stat_boxplot()`).

## Warning: Removed 1 rows containing non-finite values (`stat_boxplot()`).
## Removed 1 rows containing non-finite values (`stat_boxplot()`).
## Removed 1 rows containing non-finite values (`stat_boxplot()`).
gma year_ct site settlements subunit td_sc cos_td_rad sin_td_rad
Min. : 1.000 Min. :0.0 Nir Etzion : 31 Far :224 Judean Highlands:149 Min. :-1.9882 Min. :-0.1671 Min. :-1.0000
1st Qu.: 2.076 1st Qu.:3.0 Ein Yaakov : 30 Near:220 Carmel :150 1st Qu.:-0.2713 1st Qu.: 0.5702 1st Qu.:-0.8215
Median : 2.560 Median :5.0 Givat Yearim : 30 NA Galilee :145 Median : 0.4918 Median : 0.8140 Median :-0.5808
Mean : 2.818 Mean :4.8 Givat Yeshayahu: 30 NA NA Mean : 0.3205 Mean : 0.7098 Mean :-0.5864
3rd Qu.: 3.252 3rd Qu.:7.0 Goren : 30 NA NA 3rd Qu.: 1.1023 3rd Qu.: 0.9413 3rd Qu.:-0.3375
Max. :10.301 Max. :9.0 Kerem Maharal : 30 NA NA Max. : 1.7127 Max. : 0.9976 Max. :-0.0688
NA’s :1 NA (Other) :263 NA NA NA NA NA

Fit glm, compare gamma, gaussian (poisson inappropriate because response is not discrete)

Remove rows 211, 376, 408. Fit fixed and mixed models.

Mixed model converged

## Linear mixed model fit by REML ['lmerMod']
## Formula: gma ~ settlements * year_ct + subunit + cos_td_rad + sin_td_rad +  
##     (1 | site)
##    Data: P.anal
## 
## REML criterion at convergence: 1292.1
## 
## Scaled residuals: 
##     Min      1Q  Median      3Q     Max 
## -1.9628 -0.6076 -0.1482  0.4344  5.8495 
## 
## Random effects:
##  Groups   Name        Variance Std.Dev.
##  site     (Intercept) 0.07491  0.2737  
##  Residual             1.02960  1.0147  
## Number of obs: 440, groups:  site, 16
## 
## Fixed effects:
##                         Estimate Std. Error t value
## (Intercept)              0.99047    0.57113   1.734
## settlementsNear          0.33563    0.17729   1.893
## year_ct                 -0.01007    0.02642  -0.381
## subunitCarmel            0.13762    0.20953   0.657
## subunitGalilee          -0.07988    0.20628  -0.387
## cos_td_rad               1.17762    0.42045   2.801
## sin_td_rad              -1.48469    0.49359  -3.008
## settlementsNear:year_ct -0.02829    0.03101  -0.912
## 
## Correlation of Fixed Effects:
##             (Intr) sttlmN yer_ct sbntCr sbntGl cs_td_ sn_td_
## settlmntsNr -0.157                                          
## year_ct      0.022  0.487                                   
## subunitCrml -0.191 -0.001  0.008                            
## subunitGall -0.202  0.003  0.005  0.508                     
## cos_td_rad  -0.934  0.003 -0.106  0.012  0.015              
## sin_td_rad   0.871 -0.004  0.374  0.001 -0.014 -0.856       
## sttlmntsN:_  0.132 -0.838 -0.582  0.003  0.001 -0.004  0.003

perform stepwise model selection of gaussian model.

## Single term deletions
## 
## Model:
## gma ~ settlements * year_ct + subunit + cos_td_rad + sin_td_rad + 
##     (1 | site)
##                     npar    AIC
## <none>                   1290.5
## subunit                2 1287.8
## cos_td_rad             1 1296.3
## sin_td_rad             1 1297.7
## settlements:year_ct    1 1289.3

drop subunit

## Single term deletions
## 
## Model:
## gma ~ settlements * year_ct + cos_td_rad + sin_td_rad + (1 | 
##     site)
##                     npar    AIC
## <none>                   1287.8
## cos_td_rad             1 1293.6
## sin_td_rad             1 1295.0
## settlements:year_ct    1 1286.6

drop year X settlements

## Single term deletions
## 
## Model:
## gma ~ settlements + year_ct + cos_td_rad + sin_td_rad + (1 | 
##     site)
##             npar    AIC
## <none>           1286.6
## settlements    1 1289.0
## year_ct        1 1286.0
## cos_td_rad     1 1292.5
## sin_td_rad     1 1293.9

drop year

## Single term deletions
## 
## Model:
## gma ~ settlements + cos_td_rad + sin_td_rad + (1 | site)
##             npar    AIC
## <none>           1286.0
## settlements    1 1288.2
## cos_td_rad     1 1291.1
## sin_td_rad     1 1291.9
## Single term deletions
## 
## Model:
## gma ~ settlements + cos_td_rad + sin_td_rad + (1 | site)
##             npar    AIC
## <none>           1286.0
## settlements    1 1288.2
## cos_td_rad     1 1291.1
## sin_td_rad     1 1291.9

This is the final model:

## Linear mixed model fit by REML ['lmerMod']
## Formula: gma ~ settlements + cos_td_rad + sin_td_rad + (1 | site)
##    Data: P.anal
## 
## REML criterion at convergence: 1281.4
## 
## Scaled residuals: 
##     Min      1Q  Median      3Q     Max 
## -2.0616 -0.6198 -0.1484  0.4293  5.8897 
## 
## Random effects:
##  Groups   Name        Variance Std.Dev.
##  site     (Intercept) 0.06892  0.2625  
##  Residual             1.02964  1.0147  
## Number of obs: 440, groups:  site, 16
## 
## Fixed effects:
##                 Estimate Std. Error t value
## (Intercept)       1.1545     0.5453   2.117
## settlementsNear   0.2002     0.0968   2.069
## cos_td_rad        1.1108     0.4160   2.670
## sin_td_rad       -1.2330     0.4369  -2.823
## 
## Correlation of Fixed Effects:
##             (Intr) sttlmN cs_td_
## settlmntsNr -0.088              
## cos_td_rad  -0.964  0.000       
## sin_td_rad   0.957 -0.002 -0.904
## $site
##                  (Intercept)
## Abirim           0.008982551
## Aderet           0.378982346
## Beit Oren        0.277725813
## Ein Yaakov       0.005273928
## Givat Yearim    -0.367444219
## Givat Yeshayahu  0.158173367
## Goren            0.162039344
## Iftach          -0.305765485
## Kerem Maharal   -0.030632466
## Kfar Shamai     -0.137142196
## Margaliot       -0.074170019
## Nehusha         -0.154001749
## Nir Etzion      -0.149171842
## Ofer             0.038757806
## Ramat Raziel    -0.079372801
## Yagur            0.267765624
## 
## with conditional variances for "site"
## $site

Not a great fit. settlement is significant.

Observations 440
Dependent variable gma
Type Mixed effects linear regression
AIC 1293.430
BIC 1317.950
Pseudo-R² (fixed effects) 0.028
Pseudo-R² (total) 0.089
Fixed Effects
Est. S.E. t val. d.f. p
(Intercept) 1.155 0.549 2.102 406.354 0.036
settlementsNear 0.200 0.097 2.068 421.708 0.039
cos_td_rad 1.111 0.419 2.651 427.380 0.008
sin_td_rad -1.233 0.440 -2.800 416.685 0.005
p values calculated using Kenward-Roger standard errors and d.f.
Random Effects
Group Parameter Std. Dev.
site (Intercept) 0.263
Residual 1.015
Grouping Variables
Group # groups ICC
site 16 0.063

## Confidence intervals for merMod models is an experimental feature. The
## intervals reflect only the variance of the fixed effects, not the random
## effects.
## Warning: Removed 16 rows containing missing values (`geom_point()`).
## Removed 16 rows containing missing values (`geom_point()`).

## Warning: Removed 16 rows containing missing values (`geom_point()`).

On average, GMA in near plots is higher by 7.5128056 percent.

abundance

Explore data

## [1] "ABUNDANCE WITHOUT RARE SPECIES"
abundance year_ct site settlements subunit td_sc cos_td_rad sin_td_rad
Min. : 0.00 Min. :0.0 Nir Etzion : 31 Far :224 Judean Highlands:149 Min. :-1.9882 Min. :-0.1671 Min. :-1.0000
1st Qu.: 15.75 1st Qu.:3.0 Ein Yaakov : 30 Near:220 Carmel :150 1st Qu.:-0.2713 1st Qu.: 0.5702 1st Qu.:-0.8215
Median : 24.00 Median :5.0 Givat Yearim : 30 NA Galilee :145 Median : 0.4918 Median : 0.8140 Median :-0.5808
Mean : 30.45 Mean :4.8 Givat Yeshayahu: 30 NA NA Mean : 0.3205 Mean : 0.7098 Mean :-0.5864
3rd Qu.: 36.00 3rd Qu.:7.0 Goren : 30 NA NA 3rd Qu.: 1.1023 3rd Qu.: 0.9413 3rd Qu.:-0.3375
Max. :345.00 Max. :9.0 Kerem Maharal : 30 NA NA Max. : 1.7127 Max. : 0.9976 Max. :-0.0688
NA NA (Other) :263 NA NA NA NA NA

Some outliers with total abundance >100. Examine:

##             subunit             point_name            datetime monitors_name
## 1: Judean Highlands  Givat Yeshayahu Far 2 2017-04-20 10:20:00   Eran Banker
## 2: Judean Highlands Givat Yeshayahu Near 1 2017-04-20 06:25:00   Eran Banker
## 3: Judean Highlands Givat Yeshayahu Near 2 2017-04-20 06:50:00   Eran Banker
## 4: Judean Highlands Givat Yeshayahu Near 3 2017-04-20 07:15:00   Eran Banker
## 5: Judean Highlands          Aderet Near 2 2019-05-26 06:52:00   Eran Banker
## 6:           Carmel       Beit Oren Near 1 2019-04-19 07:49:00         Other
## 7:           Carmel       Beit Oren Near 3 2019-04-19 07:35:00         Other
## 8:           Carmel   Kerem Maharal Near 3 2021-04-17 07:41:00   Eliraz Dvir
## 9:           Carmel      Nir Etzion Far 11 2021-04-23 07:24:00   Eliraz Dvir
##    richness       gma abundance
## 1:       11  5.356635       115
## 2:       13  9.170560       140
## 3:       14  7.652753       138
## 4:       16  9.098580       221
## 5:       15  5.408017       106
## 6:        5  8.043088       129
## 7:       10  7.338970       254
## 8:       11 10.301194       345
## 9:        9  3.066833       119

Exclude 3 plots with high abundance (>150) to improve model fit.

PHI>1, hence choose negative binomial. Fit fixed and mixed models. Choose mixed model if possible, otherwise choose a model with fixed-effects only.

## [1] 3689.581
## Warning in checkConv(attr(opt, "derivs"), opt$par, ctrl = control$checkConv, :
## Model failed to converge with max|grad| = 0.004264 (tol = 0.002, component 1)
## [1] 3681.608

mixed model converged, slightly better AIC. Perform stepwise model selection of mixed model.

## Warning in checkConv(attr(opt, "derivs"), opt$par, ctrl = control$checkConv, :
## Model failed to converge with max|grad| = 0.00218393 (tol = 0.002, component 1)
## Single term deletions
## 
## Model:
## abundance ~ settlements * year_ct + subunit + cos_td_rad + sin_td_rad + 
##     (1 | site)
##                     npar    AIC
## <none>                   3681.6
## subunit                2 3687.1
## cos_td_rad             1 3697.1
## sin_td_rad             1 3699.5
## settlements:year_ct    1 3679.7

drop settlements X year because \(\Delta AIC<2\)

## Single term deletions
## 
## Model:
## abundance ~ settlements + subunit + year_ct + cos_td_rad + sin_td_rad + 
##     (1 | site)
##             npar    AIC
## <none>           3679.7
## settlements    1 3773.3
## subunit        2 3685.2
## year_ct        1 3678.2
## cos_td_rad     1 3695.2
## sin_td_rad     1 3697.6

drop year

## Single term deletions
## 
## Model:
## abundance ~ settlements + subunit + cos_td_rad + sin_td_rad + 
##     (1 | site)
##             npar    AIC
## <none>           3678.2
## settlements    1 3772.0
## subunit        2 3683.8
## cos_td_rad     1 3693.2
## sin_td_rad     1 3697.8

Subunit, settlement and time of year remain. The final model:

## Generalized linear mixed model fit by maximum likelihood (Laplace
##   Approximation) [glmerMod]
##  Family: Negative Binomial(3.3101)  ( log )
## Formula: abundance ~ settlements + subunit + cos_td_rad + sin_td_rad +  
##     (1 | site)
##    Data: P.anal
## 
##      AIC      BIC   logLik deviance df.resid 
##   3678.2   3711.0  -1831.1   3662.2      436 
## 
## Scaled residuals: 
##     Min      1Q  Median      3Q     Max 
## -1.7208 -0.7138 -0.2446  0.3842  9.2005 
## 
## Random effects:
##  Groups Name        Variance Std.Dev.
##  site   (Intercept) 0.01938  0.1392  
## Number of obs: 444, groups:  site, 16
## 
## Fixed effects:
##                 Estimate Std. Error z value Pr(>|z|)    
## (Intercept)      1.67516    0.32509   5.153 2.57e-07 ***
## settlementsNear  0.54579    0.05556   9.823  < 2e-16 ***
## subunitCarmel    0.16956    0.11089   1.529   0.1262    
## subunitGalilee  -0.22714    0.11015  -2.062   0.0392 *  
## cos_td_rad       1.02590    0.24294   4.223 2.41e-05 ***
## sin_td_rad      -1.16744    0.24800  -4.708 2.51e-06 ***
## ---
## Signif. codes:  0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1
## 
## Correlation of Fixed Effects:
##             (Intr) sttlmN sbntCr sbntGl cs_td_
## settlmntsNr -0.111                            
## subunitCrml -0.214  0.020                     
## subunitGall -0.226  0.033  0.505              
## cos_td_rad  -0.946  0.019  0.050  0.043       
## sin_td_rad   0.938 -0.017 -0.032 -0.062 -0.903
## $site
##                 (Intercept)
## Abirim          -0.11137155
## Aderet           0.16235427
## Beit Oren        0.11023799
## Ein Yaakov       0.01625407
## Givat Yearim    -0.19886986
## Givat Yeshayahu  0.17665857
## Goren           -0.01370126
## Iftach           0.07733150
## Kerem Maharal    0.13673950
## Kfar Shamai      0.00869344
## Margaliot        0.02457356
## Nehusha         -0.07214524
## Nir Etzion      -0.15864707
## Ofer            -0.04190665
## Ramat Raziel    -0.06677795
## Yagur           -0.04538202
## 
## with conditional variances for "site"
## $site

Not a great fit, high residuals. center time of year variables, highly correlated with intercept.

## Generalized linear mixed model fit by maximum likelihood (Laplace
##   Approximation) [glmerMod]
##  Family: Negative Binomial(3.3101)  ( log )
## Formula: abundance ~ settlements + subunit + cos_td_rad_c + sin_td_rad_c +  
##     (1 | site)
##    Data: P.anal
## 
##      AIC      BIC   logLik deviance df.resid 
##   3678.2   3711.0  -1831.1   3662.2      436 
## 
## Scaled residuals: 
##     Min      1Q  Median      3Q     Max 
## -1.7208 -0.7138 -0.2446  0.3842  9.2005 
## 
## Random effects:
##  Groups Name        Variance Std.Dev.
##  site   (Intercept) 0.01938  0.1392  
## Number of obs: 444, groups:  site, 16
## 
## Fixed effects:
##                 Estimate Std. Error z value Pr(>|z|)    
## (Intercept)      3.08791    0.08411  36.711  < 2e-16 ***
## settlementsNear  0.54578    0.05556   9.823  < 2e-16 ***
## subunitCarmel    0.16956    0.11088   1.529   0.1262    
## subunitGalilee  -0.22714    0.11015  -2.062   0.0392 *  
## cos_td_rad_c     1.02590    0.24293   4.223 2.41e-05 ***
## sin_td_rad_c    -1.16743    0.24798  -4.708 2.51e-06 ***
## ---
## Signif. codes:  0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1
## 
## Correlation of Fixed Effects:
##             (Intr) sttlmN sbntCr sbntGl cs_t__
## settlmntsNr -0.358                            
## subunitCrml -0.669  0.020                     
## subunitGall -0.678  0.033  0.505              
## cos_td_rd_c -0.047  0.019  0.050  0.043       
## sin_td_rd_c  0.044 -0.017 -0.032 -0.062 -0.903

Interpretation of abundance model:

Observations 444
Dependent variable abundance
Type Mixed effects generalized linear model
Family Negative Binomial(3.3101)
Link log
AIC 3678.232
BIC 3710.998
Pseudo-R² (fixed effects) 0.692
Pseudo-R² (total) 0.804
Fixed Effects
exp(Est.) S.E. z val. p
(Intercept) 21.931 0.084 36.711 0.000
settlementsNear 1.726 0.056 9.823 0.000
subunitCarmel 1.185 0.111 1.529 0.126
subunitGalilee 0.797 0.110 -2.062 0.039
cos_td_rad_c 2.790 0.243 4.223 0.000
sin_td_rad_c 0.311 0.248 -4.708 0.000
Random Effects
Group Parameter Std. Dev.
site (Intercept) 0.139
Grouping Variables
Group # groups ICC
site 16 0.052
## Confidence intervals for merMod models is an experimental feature. The
## intervals reflect only the variance of the fixed effects, not the random
## effects.

## Confidence intervals for merMod models is an experimental feature. The
## intervals reflect only the variance of the fixed effects, not the random
## effects.

##        1        2 
## 21.93111 37.85224
##       2 
## 72.5961
## Confidence intervals for merMod models is an experimental feature. The
## intervals reflect only the variance of the fixed effects, not the random
## effects.

##  subunit          emmean     SE  df asymp.LCL asymp.UCL
##  Judean Highlands   3.36 0.0786 Inf      3.21      3.51
##  Carmel             3.53 0.0782 Inf      3.38      3.68
##  Galilee            3.13 0.0771 Inf      2.98      3.28
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## Confidence level used: 0.95
## [1] 49.18247
## [1] 18.53049
## [1] 25.86
##  contrast                   estimate    SE  df z.ratio p.value
##  Judean Highlands - Carmel    -0.170 0.111 Inf  -1.529  0.1262
##  Judean Highlands - Galilee    0.227 0.110 Inf   2.062  0.0588
##  Carmel - Galilee              0.397 0.110 Inf   3.607  0.0009
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## P value adjustment: fdr method for 3 tests

significantly lower abundance in Galilee subunit and far from settlements. No significant temporal trend.

community analysis using package MVabund

## Overlapping points were shifted along the y-axis to make them visible.
## 
##  PIPING TO 2nd MVFACTOR
## Only the variables Curruca.melanocephala, Turdus.merula, Streptopelia.decaocto, Passer.domesticus, Pycnonotus.xanthopygos, Columba.livia, Parus.major, Garrulus.glandarius, Cinnyris.osea, Spilopelia.senegalensis, Cecropis.daurica, Chloris.chloris were included in the plot 
## (the variables with highest total abundance).

## Overlapping points were shifted along the y-axis to make them visible.
## 
##  PIPING TO 2nd MVFACTOR
## Only the variables Curruca.melanocephala, Turdus.merula, Streptopelia.decaocto, Passer.domesticus, Pycnonotus.xanthopygos, Columba.livia, Parus.major, Garrulus.glandarius, Cinnyris.osea, Spilopelia.senegalensis, Cecropis.daurica, Chloris.chloris were included in the plot 
## (the variables with highest total abundance).

There are few observations with counts of >60. Examine these:

##              point_name            datetime             SciName monitors_name
## 1: Kerem Maharal Near 3 2021-04-17 07:41:00     Chloris chloris   Eliraz Dvir
## 2: Kerem Maharal Near 3 2021-04-17 07:41:00 Garrulus glandarius   Eliraz Dvir
## 3:     Beit Oren Near 3 2019-04-19 07:35:00     Curruca curruca         Other
## 4:     Beit Oren Near 1 2019-04-19 07:49:00     Curruca curruca         Other
##    notes rad_0_20 rad_20_100 rad_100_250 rad_over_250
## 1:              0         80           0            0
## 2:              0          0         150            0
## 3:  <NA>       12         30         100            0
## 4:  <NA>       14         35          50            0

G. glandarius observation seems highly irregular. Remove altogether, might be a mistake. C. curruca observations are probably due to migration waves. Remove.

start model specification:

##       nb       po 
##  978.686 1390.834
## [1] "POISSON"

## [1] "NEGATIVE BINOMIAL"

negative binomial model is better than poisson according to residuals and AIC comparison.

##        nb        po       nb1 
##  978.6860 1390.8344  968.2392

The addition of the explanatory variable ‘site’ is somewhat improving the AIC of the model. Prefer to exclude site, for simplification. stepwise selection of model:

## Single term deletions
## 
## Model:
## spp_no_rare ~ settlements * year_ct + subunit + cos_td_rad + 
##     sin_td_rad
##                     Df   AIC
## <none>                 23489
## subunit             48 23844
## cos_td_rad          24 23537
## sin_td_rad          24 23549
## settlements:year_ct 24 23481

drop settlements X year.

## Single term deletions
## 
## Model:
## spp_no_rare ~ settlements + subunit + year_ct + cos_td_rad + 
##     sin_td_rad
##             Df   AIC
## <none>         23481
## settlements 24 24269
## subunit     48 23838
## year_ct     24 23502
## cos_td_rad  24 23528
## sin_td_rad  24 23542

final model includes settlements, year, subunit, sampling time of year.

## 
## Test statistics:
##                 wald value Pr(>wald)    
## (Intercept)         12.055     0.001 ***
## settlementsNear     28.826     0.001 ***
## subunitCarmel       14.027     0.001 ***
## subunitGalilee      15.305     0.001 ***
## year_ct              8.913     0.001 ***
## cos_td_rad          10.020     0.001 ***
## sin_td_rad          10.820     0.001 ***
## --- 
## Signif. codes:  0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1 
## 
## Test statistic:  38.38, p-value: 0.001 
## Arguments:
##  Test statistics calculated assuming response assumed to be uncorrelated 
##  P-value calculated using 999 resampling iterations via pit.trap resampling (to account for correlation in testing).
## Analysis of Deviance Table
## 
## Model: spp_no_rare ~ settlements + subunit + year_ct + cos_td_rad + sin_td_rad
## 
## Multivariate test:
##             Res.Df Df.diff   Dev Pr(>Dev)   
## (Intercept)    443                          
## settlements    442       1 770.4     0.01 **
## subunit        440       2 426.6     0.01 **
## year_ct        439       1 143.4     0.01 **
## cos_td_rad     438       1  46.9     0.04 * 
## sin_td_rad     437       1 109.1     0.01 **
## ---
## Signif. codes:  0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1
## 
## Univariate Tests:
##             Acridotheres.tristis          Alectoris.chukar         
##                              Dev Pr(>Dev)              Dev Pr(>Dev)
## (Intercept)                                                        
## settlements               79.707     0.01           38.366     0.01
## subunit                   39.138     0.01           10.607     0.10
## year_ct                   29.844     0.01            9.298     0.07
## cos_td_rad                 3.002     0.87            2.276     0.95
## sin_td_rad                  5.64     0.41            0.596     1.00
##             Carduelis.carduelis          Cecropis.daurica         
##                             Dev Pr(>Dev)              Dev Pr(>Dev)
## (Intercept)                                                       
## settlements                8.13     0.04           33.414     0.01
## subunit                  21.019     0.01            1.066     0.94
## year_ct                   6.527     0.29            1.257     0.96
## cos_td_rad                0.389     1.00            0.127     1.00
## sin_td_rad                 1.61     0.95            3.044     0.78
##             Chloris.chloris          Cinnyris.osea          Columba.livia
##                         Dev Pr(>Dev)           Dev Pr(>Dev)           Dev
## (Intercept)                                                              
## settlements           30.55     0.01        41.948     0.01         6.276
## subunit               0.613     0.94        17.867     0.02        19.317
## year_ct               5.072     0.46         3.939     0.62        20.916
## cos_td_rad            0.072     1.00         1.651     0.98          0.01
## sin_td_rad            0.112     1.00          2.01     0.93        11.564
##                      Corvus.cornix          Corvus.monedula         
##             Pr(>Dev)           Dev Pr(>Dev)             Dev Pr(>Dev)
## (Intercept)                                                         
## settlements     0.11        22.932     0.01           0.428     0.58
## subunit         0.01        47.477     0.01          22.714     0.01
## year_ct         0.01         0.695     0.98           0.301     1.00
## cos_td_rad      1.00         1.013     1.00           0.184     1.00
## sin_td_rad      0.02         0.157     1.00           0.053     1.00
##             Curruca.curruca          Curruca.melanocephala         
##                         Dev Pr(>Dev)                   Dev Pr(>Dev)
## (Intercept)                                                        
## settlements           2.096     0.58                 21.24     0.01
## subunit              27.501     0.01                41.067     0.01
## year_ct              13.288     0.02                 0.747     0.98
## cos_td_rad             7.65     0.23                   0.3     1.00
## sin_td_rad            4.442     0.56                 0.429     1.00
##             Dendrocopos.syriacus          Falco.tinnunculus         
##                              Dev Pr(>Dev)               Dev Pr(>Dev)
## (Intercept)                                                         
## settlements                9.285     0.03             1.849     0.58
## subunit                    3.648     0.66             9.122     0.11
## year_ct                    0.002     1.00             0.543     0.98
## cos_td_rad                 1.417     1.00             0.078     1.00
## sin_td_rad                  3.35     0.71             0.641     1.00
##             Garrulus.glandarius          Parus.major          Passer.domesticus
##                             Dev Pr(>Dev)         Dev Pr(>Dev)               Dev
## (Intercept)                                                                    
## settlements               3.502     0.44       3.908     0.44           166.539
## subunit                  24.625     0.01      25.606     0.01            21.305
## year_ct                   4.924     0.46       0.176     1.00             1.781
## cos_td_rad                1.156     1.00       3.376     0.84             0.485
## sin_td_rad                1.081     0.99       7.701     0.17             2.962
##                      Prinia.gracilis          Psittacula.krameri         
##             Pr(>Dev)             Dev Pr(>Dev)                Dev Pr(>Dev)
## (Intercept)                                                              
## settlements     0.01          48.787     0.01             15.594     0.01
## subunit         0.01           1.864     0.90             14.393     0.05
## year_ct         0.92          16.036     0.02              1.082     0.96
## cos_td_rad      1.00           0.633     1.00              0.793     1.00
## sin_td_rad      0.80           1.004     0.99              1.724     0.95
##             Pycnonotus.xanthopygos          Spilopelia.senegalensis         
##                                Dev Pr(>Dev)                     Dev Pr(>Dev)
## (Intercept)                                                                 
## settlements                 45.339     0.01                 157.013     0.01
## subunit                      25.29     0.01                  27.046     0.01
## year_ct                       2.87     0.78                   2.224     0.88
## cos_td_rad                  11.394     0.05                   0.333     1.00
## sin_td_rad                   0.499     1.00                   0.603     1.00
##             Streptopelia.decaocto          Streptopelia.turtur         
##                               Dev Pr(>Dev)                 Dev Pr(>Dev)
## (Intercept)                                                            
## settlements                 1.955     0.58               1.175     0.58
## subunit                    12.168     0.09               6.566     0.33
## year_ct                     0.068     1.00              18.089     0.02
## cos_td_rad                  6.672     0.34               3.045     0.87
## sin_td_rad                 14.008     0.01               1.847     0.93
##             Troglodytes.troglodytes          Turdus.merula         
##                                 Dev Pr(>Dev)           Dev Pr(>Dev)
## (Intercept)                                                        
## settlements                  26.472     0.01         3.926     0.44
## subunit                       4.411     0.62         2.142     0.90
## year_ct                       0.576     0.98         3.137     0.75
## cos_td_rad                     0.48     1.00         0.347     1.00
## sin_td_rad                   10.786     0.03         33.19     0.01
## Arguments:
##  Test statistics calculated assuming uncorrelated response (for faster computation) 
## P-value calculated using 99 iterations via PIT-trap resampling.

Factors settlements, year, subunit, time of year have a statistically significant effect on community composition.

## Loading required package: ggnewscale
## Warning: Using `size` aesthetic for lines was deprecated in ggplot2 3.4.0.
## ℹ Please use `linewidth` instead.
## This warning is displayed once every 8 hours.
## Call `lifecycle::last_lifecycle_warnings()` to see where this warning was
## generated.
## Warning: The `size` argument of `element_line()` is deprecated as of ggplot2 3.4.0.
## ℹ Please use the `linewidth` argument instead.
## This warning is displayed once every 8 hours.
## Call `lifecycle::last_lifecycle_warnings()` to see where this warning was
## generated.
## [1] "black arrow is median of synanthrope/invasive\n\n"

## [1] "black arrow is median of synanthrope/invasive\n\n"

## [[1]]

## 
## [[2]]

## 
## [[3]]

Alectoris chukar - חוגלת סלעים

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##      -3.3297204      -1.8142174       1.0363737       0.3667818       0.1915418 
##      cos_td_rad      sin_td_rad 
##       1.7562492      -1.0706266
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##      -3.3297134      -1.8142178       1.0363750       0.3667830       0.1915409 
##      cos_td_rad      sin_td_rad 
##       1.7562446      -1.0706260

## [1] 460.6174
##   [1] 0.2333147 0.2374129 0.2415831 0.2458266 0.2501446 0.2545385 0.2590095
##   [8] 0.2635591 0.2681886 0.2728994 0.2776930 0.2825708 0.2875342 0.2925848
##  [15] 0.2977242 0.3029538 0.3082752 0.3136902 0.3192003 0.3248071 0.3305124
##  [22] 0.3363180 0.3422255 0.3482368 0.3543537 0.3605780 0.3669117 0.3733566
##  [29] 0.3799147 0.3865880 0.3933786 0.4002884 0.4073196 0.4144743 0.4217546
##  [36] 0.4291629 0.4367012 0.4443720 0.4521776 0.4601202 0.4682023 0.4764265
##  [43] 0.4847950 0.4933106 0.5019758 0.5107931 0.5197653 0.5288952 0.5381854
##  [50] 0.5476388 0.5572582 0.5670466 0.5770070 0.5871423 0.5974556 0.6079501
##  [57] 0.6186289 0.6294953 0.6405526 0.6518041 0.6632532 0.6749035 0.6867583
##  [64] 0.6988215 0.7110965 0.7235871 0.7362971 0.7492304 0.7623908 0.7757825
##  [71] 0.7894093 0.8032755 0.8173853 0.8317429 0.8463527 0.8612192 0.8763468
##  [78] 0.8917400 0.9074037 0.9233425 0.9395613 0.9560650 0.9728586 0.9899471
##  [85] 1.0073359 1.0250300 1.0430350 1.0613562 1.0799992 1.0989697 1.1182735
##  [92] 1.1379163 1.1579041 1.1782431 1.1989393 1.2199990 1.2414286 1.2632347
##  [99] 1.2854238 1.3080027
## [1] 4.606174

## [1] 513.6274
## [1] 5.136275
## [1] 5.887509

Acridotheres tristis - מיינה מצויה

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##     -5.81322891      3.04949199     -0.06360271     -2.84839762      0.14854946 
##      cos_td_rad      sin_td_rad 
##      1.04586057     -2.82696864
##     (Intercept) settlementsNear         year_ct   subunitCarmel  subunitGalilee 
##     -5.81322905      3.04949186      0.14854959     -0.06360287     -2.84839757 
##      cos_td_rad      sin_td_rad 
##      1.04586061     -2.82696797

## [1] 280.7399
##   [1] 0.03293719 0.03338501 0.03383892 0.03429899 0.03476533 0.03523800
##   [7] 0.03571710 0.03620271 0.03669493 0.03719384 0.03769953 0.03821209
##  [13] 0.03873163 0.03925823 0.03979199 0.04033300 0.04088138 0.04143720
##  [19] 0.04200059 0.04257163 0.04315044 0.04373712 0.04433177 0.04493451
##  [25] 0.04554545 0.04616469 0.04679235 0.04742854 0.04807338 0.04872700
##  [31] 0.04938949 0.05006100 0.05074163 0.05143152 0.05213079 0.05283956
##  [37] 0.05355798 0.05428616 0.05502424 0.05577235 0.05653064 0.05729924
##  [43] 0.05807828 0.05886792 0.05966830 0.06047955 0.06130184 0.06213530
##  [49] 0.06298010 0.06383639 0.06470431 0.06558404 0.06647573 0.06737954
##  [55] 0.06829564 0.06922419 0.07016537 0.07111935 0.07208629 0.07306639
##  [61] 0.07405981 0.07506673 0.07608735 0.07712184 0.07817039 0.07923321
##  [67] 0.08031047 0.08140238 0.08250913 0.08363094 0.08476799 0.08592051
##  [73] 0.08708869 0.08827276 0.08947292 0.09068941 0.09192243 0.09317222
##  [79] 0.09443900 0.09572300 0.09702446 0.09834361 0.09968070 0.10103597
##  [85] 0.10240967 0.10380204 0.10521335 0.10664384 0.10809378 0.10956343
##  [91] 0.11105307 0.11256296 0.11409338 0.11564460 0.11721692 0.11881061
##  [97] 0.12042597 0.12206330 0.12372288 0.12540503
## [1] 2.807399
## [1] 3.160956

## [1] 2010.462
## [1] 0.0671933 1.4180889
## [1] 20.10462
## [1] 21.10462
##  subunit          emmean    SE  df asymp.LCL asymp.UCL
##  Judean Highlands  -1.18 0.216 Inf     -1.60    -0.753
##  Carmel            -1.24 0.220 Inf     -1.67    -0.808
##  Galilee           -4.02 0.430 Inf     -4.87    -3.180
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## Confidence level used: 0.95
## [1] 1511.902
## [1] 6.183655
## [1] 1611.577
##  contrast                   estimate    SE  df z.ratio p.value
##  Judean Highlands - Carmel    0.0636 0.255 Inf   0.249  0.8032
##  Judean Highlands - Galilee   2.8484 0.435 Inf   6.547  <.0001
##  Carmel - Galilee             2.7848 0.433 Inf   6.436  <.0001
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## P value adjustment: fdr method for 3 tests

Carduelis carduelis - חוחית

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##      -6.3665442       1.2522516       2.4837754       0.9127661      -0.1859762 
##      cos_td_rad      sin_td_rad 
##       2.5842809      -2.6428473
##     (Intercept) settlementsNear         year_ct   subunitCarmel  subunitGalilee 
##      -6.3665485       1.2522511      -0.1859760       2.4837743       0.9127662 
##      cos_td_rad      sin_td_rad 
##       2.5842828      -2.6428525

## [1] 249.8209
## [1] 2.498209

##  subunit          emmean    SE  df asymp.LCL asymp.UCL
##  Judean Highlands -3.249 0.457 Inf     -4.15    -2.352
##  Carmel           -0.765 0.259 Inf     -1.27    -0.258
##  Galilee          -2.336 0.354 Inf     -3.03    -1.642
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## Confidence level used: 0.95
## [1] 381.1457
## [1] 1098.913
## [1] 149.1787
##  contrast                   estimate    SE  df z.ratio p.value
##  Judean Highlands - Carmel    -2.484 0.516 Inf  -4.810  <.0001
##  Judean Highlands - Galilee   -0.913 0.561 Inf  -1.627  0.1038
##  Carmel - Galilee              1.571 0.432 Inf   3.635  0.0004
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## P value adjustment: fdr method for 3 tests

Cecropis daurica - סנונית מערות

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##      0.98095288      2.16869649     -0.45368366     -0.08874088      0.10836920 
##      cos_td_rad      sin_td_rad 
##     -2.15241967      2.63508635
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##      0.98094089      2.16869257     -0.45368117     -0.08874169      0.10836879 
##      sin_td_rad      cos_td_rad 
##      2.63507576     -2.15240660

## [1] 774.6841
## [1] 0.2076556 1.8163309
## [1] 7.746843

Chloris chloris - ירקון

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##     -1.17270658      1.46825586     -0.42961211     -0.09156341      0.11302623 
##      cos_td_rad      sin_td_rad 
##     -0.19368715      0.42817036
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##     -1.17266422      1.46823977     -0.42962756     -0.09155752      0.11302354 
##      sin_td_rad      cos_td_rad 
##      0.42819754     -0.19369059

## [1] 334.1586
## [1] 0.3610366 1.5674714
## [1] 3.341586

Cinnyris osea - צופית בוהקת

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##     -0.51622489      0.87058980     -0.26514518     -0.71725657     -0.07193677 
##      cos_td_rad      sin_td_rad 
##      0.27528358     -0.82378562
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##     -0.51622477      0.87058991     -0.26514518     -0.71725663     -0.07193676 
##      cos_td_rad      sin_td_rad 
##      0.27528327     -0.82378562

## [1] 138.8319
## [1] 0.832746 1.988863
## [1] 1.388319

##  subunit           emmean    SE  df asymp.LCL asymp.UCL
##  Judean Highlands  0.2523 0.103 Inf    0.0507     0.454
##  Carmel           -0.0129 0.110 Inf   -0.2277     0.202
##  Galilee          -0.4650 0.128 Inf   -0.7152    -0.215
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## Confidence level used: 0.95
## [1] 57.16091
## [1] 30.36917
## [1] 104.8894
##  contrast                   estimate    SE  df z.ratio p.value
##  Judean Highlands - Carmel     0.265 0.149 Inf   1.785  0.0743
##  Judean Highlands - Galilee    0.717 0.162 Inf   4.420  <.0001
##  Carmel - Galilee              0.452 0.167 Inf   2.715  0.0100
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## P value adjustment: fdr method for 3 tests

Columba livia - יונת סלעים

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##      -8.7541198       1.8964697       0.5402957      -1.5045681       0.1794411 
##      cos_td_rad      sin_td_rad 
##       5.1582629      -5.7721640
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##      -8.7540662       1.8964544       0.5403010      -1.5045420       0.1794368 
##      cos_td_rad      sin_td_rad 
##       5.1582320      -5.7721438

## [1] 402.7542
##   [1] 0.1812109 0.1841911 0.1872204 0.1902995 0.1934292 0.1966103 0.1998438
##   [8] 0.2031305 0.2064712 0.2098669 0.2133184 0.2168267 0.2203927 0.2240173
##  [15] 0.2277015 0.2314464 0.2352528 0.2391218 0.2430544 0.2470518 0.2511148
##  [22] 0.2552447 0.2594425 0.2637094 0.2680464 0.2724548 0.2769356 0.2814901
##  [29] 0.2861196 0.2908252 0.2956081 0.3004698 0.3054114 0.3104342 0.3155397
##  [36] 0.3207291 0.3260039 0.3313654 0.3368151 0.3423545 0.3479849 0.3537079
##  [43] 0.3595251 0.3654379 0.3714480 0.3775569 0.3837663 0.3900778 0.3964931
##  [50] 0.4030139 0.4096419 0.4163790 0.4232269 0.4301873 0.4372623 0.4444536
##  [57] 0.4517632 0.4591930 0.4667450 0.4744211 0.4822236 0.4901543 0.4982155
##  [64] 0.5064093 0.5147378 0.5232033 0.5318080 0.5405542 0.5494443 0.5584805
##  [71] 0.5676654 0.5770014 0.5864909 0.5961364 0.6059406 0.6159060 0.6260353
##  [78] 0.6363313 0.6467965 0.6574338 0.6682461 0.6792363 0.6904071 0.7017617
##  [85] 0.7133030 0.7250341 0.7369582 0.7490784 0.7613979 0.7739200 0.7866480
##  [92] 0.7995854 0.8127356 0.8261020 0.8396882 0.8534979 0.8675347 0.8818024
##  [99] 0.8963047 0.9110455
## [1] 4.027542

##  subunit          emmean    SE  df asymp.LCL asymp.UCL
##  Judean Highlands  0.101 0.254 Inf    -0.397     0.600
##  Carmel            0.642 0.247 Inf     0.157     1.127
##  Galilee          -1.403 0.307 Inf    -2.005    -0.801
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## Confidence level used: 0.95
## [1] 672.9159
## [1] 71.77237
## [1] 349.9652
##  contrast                   estimate    SE  df z.ratio p.value
##  Judean Highlands - Carmel     -0.54 0.354 Inf  -1.528  0.1265
##  Judean Highlands - Galilee     1.50 0.394 Inf   3.815  0.0002
##  Carmel - Galilee               2.04 0.390 Inf   5.243  <.0001
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## P value adjustment: fdr method for 3 tests

Corvus cornix - עורב אפור

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##    -2.033847764     1.253931156     1.716084872     0.856113981     0.003008888 
##      cos_td_rad      sin_td_rad 
##     0.134731455     0.386707210
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##    -2.033846968     1.253930552     1.716084018     0.856113449     0.003008826 
##      cos_td_rad      sin_td_rad 
##     0.134731422     0.386706516

## [1] 250.4089
## [1] 0.1164205 0.4079478
## [1] 2.504089
## [1] 3.504089

##  subunit          emmean    SE  df asymp.LCL asymp.UCL
##  Judean Highlands -1.524 0.213 Inf   -1.9408    -1.106
##  Carmel            0.193 0.146 Inf   -0.0927     0.478
##  Galilee          -0.667 0.173 Inf   -1.0065    -0.328
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## Confidence level used: 0.95
## [1] 136.3161
## [1] 456.78
## [1] 135.6082
##  contrast                   estimate    SE  df z.ratio p.value
##  Judean Highlands - Carmel    -1.716 0.255 Inf  -6.724  <.0001
##  Judean Highlands - Galilee   -0.856 0.270 Inf  -3.168  0.0015
##  Carmel - Galilee              0.860 0.224 Inf   3.838  0.0002
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## P value adjustment: fdr method for 3 tests

Curruca curruca - סבכי טוחנים

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##    -19.51596678     -0.76748669     11.66738622     12.74590976      0.06434301 
##      cos_td_rad      sin_td_rad 
##      1.31234280     -6.42805839
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##    -41.73709897     -0.76749621     33.88839482     34.96693121      0.06434496 
##      cos_td_rad      sin_td_rad 
##      1.31237217     -6.42817587
## Error in qr.solve(qr.R(qr.lm(object))[p1, p1]): singular matrix 'a' in solve
## Error in eval(expr, envir, enclos): object 'curruca_time' not found
## Error in eval(expr, envir, enclos): object 'curruca_time' not found
## Error in eval(expr, envir, enclos): object 'curruca_time' not found
## [1] 4.027542
## [1] 5.027542
## Error in qr.solve(qr.R(qr.lm(object))[p1, p1]): singular matrix 'a' in solve
## Error in eval(expr, envir, enclos): object 'curruca_subunit' not found
##  subunit          emmean      SE  df asymp.LCL asymp.UCL
##  Judean Highlands -37.11 4717424 Inf  -9246019   9245945
##  Carmel            -3.22       0 Inf        -4        -2
##  Galilee           -2.14       0 Inf        -3        -1
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## Confidence level used: 0.95
## [1] 194.468
## [1] 5.332439e+16
## [1] 1.570232e+17
##  contrast                   estimate      SE  df z.ratio p.value
##  Judean Highlands - Carmel    -33.89 4717424 Inf   0.000  1.0000
##  Judean Highlands - Galilee   -34.97 4717424 Inf   0.000  1.0000
##  Carmel - Galilee              -1.08       0 Inf  -2.306  0.0633
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## P value adjustment: fdr method for 3 tests

Curruca melanocephala - סבכי שחור-ראש

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##      1.62651788     -0.42701053     -0.06039957     -0.66627444     -0.01213706 
##      cos_td_rad      sin_td_rad 
##      0.29890567     -0.26265011
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##      1.62651794     -0.42701053     -0.06039957     -0.66627442     -0.01213706 
##      cos_td_rad      sin_td_rad 
##      0.29890565     -0.26265002

## [1] -0.5326688
## [1] 53.26688
## [1] 1.532669

##  subunit          emmean     SE  df asymp.LCL asymp.UCL
##  Judean Highlands   1.72 0.0713 Inf     1.581      1.86
##  Carmel             1.66 0.0716 Inf     1.520      1.80
##  Galilee            1.05 0.0799 Inf     0.898      1.21
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## Confidence level used: 0.95
## [1] 84.04314
## [1] 6.183655
## [1] 95.42373
##  contrast                   estimate    SE  df z.ratio p.value
##  Judean Highlands - Carmel    0.0604 0.101 Inf   0.598  0.5499
##  Judean Highlands - Galilee   0.6663 0.107 Inf   6.229  <.0001
##  Carmel - Galilee             0.6059 0.107 Inf   5.653  <.0001
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## P value adjustment: fdr method for 3 tests

Dendrocopos syriacus - נקר סורי

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##    -6.572022180     0.934448120    -0.650716218    -0.238886114    -0.004452597 
##      cos_td_rad      sin_td_rad 
##     3.295442948    -2.857773539
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##    -6.572021986     0.934448131    -0.650716326    -0.238886302    -0.004452595 
##      cos_td_rad      sin_td_rad 
##     3.295442934    -2.857773352

## [1] 154.5808
## [1] 0.07588191 0.19318078
## [1] 1.545808
## [1] 2.545808

Passer domesticus - דרור הבית

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##      -5.2174230       4.9808727      -1.2576557       0.3878957      -0.1176087 
##      cos_td_rad      sin_td_rad 
##       1.4382456      -2.3168731
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##      -5.2174237       4.9808727      -1.2576557       0.3878958      -0.1176087 
##      cos_td_rad      sin_td_rad 
##       1.4382462      -2.3168737

## [1] 14460.14
## [1] 144.6014
## [1] 156.3748

##  subunit          emmean    SE  df asymp.LCL asymp.UCL
##  Judean Highlands -0.912 0.270 Inf     -1.44   -0.3838
##  Carmel           -2.170 0.304 Inf     -2.76   -1.5744
##  Galilee          -0.524 0.260 Inf     -1.03   -0.0151
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## Confidence level used: 0.95
## [1] 418.6194
## [1] 251.8378
## [1] 47.40298
##  contrast                   estimate    SE  df z.ratio p.value
##  Judean Highlands - Carmel     1.258 0.313 Inf   4.022  0.0001
##  Judean Highlands - Galilee   -0.388 0.296 Inf  -1.311  0.1899
##  Carmel - Galilee             -1.646 0.315 Inf  -5.230  <.0001
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## P value adjustment: fdr method for 3 tests

Prinia gracilis - פשוש

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##     -0.12702277      1.51717609      0.26228731     -0.06852479     -0.08504256 
##      cos_td_rad      sin_td_rad 
##     -0.36574149      0.92278481
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##     -0.12702190      1.51717688      0.26228733     -0.06852537     -0.08504253 
##      cos_td_rad      sin_td_rad 
##     -0.36574233      0.92278602

## [1] 355.9335
## [1] 0.262919 1.198736
## [1] 3.559336

## [1] 0.5348442
##   [1] 0.3954450 0.3923996 0.3893776 0.3863788 0.3834032 0.3804505 0.3775205
##   [8] 0.3746131 0.3717281 0.3688653 0.3660245 0.3632057 0.3604085 0.3576329
##  [15] 0.3548786 0.3521456 0.3494336 0.3467425 0.3440721 0.3414223 0.3387929
##  [22] 0.3361837 0.3335947 0.3310255 0.3284762 0.3259465 0.3234363 0.3209454
##  [29] 0.3184737 0.3160210 0.3135872 0.3111722 0.3087758 0.3063978 0.3040381
##  [36] 0.3016966 0.2993731 0.2970676 0.2947797 0.2925096 0.2902568 0.2880215
##  [43] 0.2858033 0.2836023 0.2814181 0.2792509 0.2771002 0.2749662 0.2728486
##  [50] 0.2707473 0.2686622 0.2665931 0.2645400 0.2625027 0.2604811 0.2584750
##  [57] 0.2564844 0.2545092 0.2525491 0.2506041 0.2486741 0.2467590 0.2448587
##  [64] 0.2429729 0.2411017 0.2392449 0.2374024 0.2355741 0.2337598 0.2319596
##  [71] 0.2301732 0.2284005 0.2266416 0.2248961 0.2231641 0.2214455 0.2197400
##  [78] 0.2180477 0.2163685 0.2147022 0.2130487 0.2114079 0.2097798 0.2081642
##  [85] 0.2065611 0.2049703 0.2033917 0.2018253 0.2002710 0.1987287 0.1971982
##  [92] 0.1956795 0.1941725 0.1926771 0.1911932 0.1897208 0.1882597 0.1868098
##  [99] 0.1853712 0.1839436
## [1] 0.534844

Psittacula krameri - דררה

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##    -5.269088823     1.668902826     1.190144622    -0.452394245    -0.009682361 
##      cos_td_rad      sin_td_rad 
##     0.995954283    -2.204367675
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##    -5.269099503     1.668904042     1.190146788    -0.452393130    -0.009682248 
##      cos_td_rad      sin_td_rad 
##     0.995960536    -2.204373874

## [1] 430.6349
## [1] 0.03629751 0.19260726
## [1] 4.306349
## [1] 5.306349

##  subunit          emmean    SE  df asymp.LCL asymp.UCL
##  Judean Highlands  -2.48 0.347 Inf     -3.16    -1.801
##  Carmel            -1.29 0.263 Inf     -1.81    -0.777
##  Galilee           -2.93 0.404 Inf     -3.72    -2.143
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## Confidence level used: 0.95
## [1] 415.517
## [1] 228.7081
## [1] 56.83122
##  contrast                   estimate    SE  df z.ratio p.value
##  Judean Highlands - Carmel    -1.190 0.419 Inf  -2.839  0.0068
##  Judean Highlands - Galilee    0.452 0.510 Inf   0.887  0.3750
##  Carmel - Galilee              1.643 0.465 Inf   3.533  0.0012
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## P value adjustment: fdr method for 3 tests

Pycnonotus.xanthopygos - בולבול ממושקף

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##    -0.887826774     1.006461173    -0.586201462    -0.823030493    -0.002510203 
##      cos_td_rad      sin_td_rad 
##     1.409560683    -0.456734212
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##    -0.887826608     1.006461068    -0.586201536    -0.823030497    -0.002510177 
##      cos_td_rad      sin_td_rad 
##     1.409560652    -0.456733882

## [1] 173.5902
## [1] 1.445443 3.954589
## [1] 1.735901
## [1] 2.735901

##  subunit          emmean    SE  df asymp.LCL asymp.UCL
##  Judean Highlands 0.8716 0.111 Inf    0.6546     1.089
##  Carmel           0.2854 0.120 Inf    0.0497     0.521
##  Galilee          0.0486 0.128 Inf   -0.2018     0.299
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## Confidence level used: 0.95
## [1] 26.71877
## [1] 79.71463
## [1] 127.7322
##  contrast                   estimate    SE  df z.ratio p.value
##  Judean Highlands - Carmel     0.586 0.163 Inf   3.605  0.0005
##  Judean Highlands - Galilee    0.823 0.168 Inf   4.899  <.0001
##  Carmel - Galilee              0.237 0.174 Inf   1.357  0.1746
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## P value adjustment: fdr method for 3 tests

Spilopelia senegalensis - צוצלת

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##     -2.48717563      2.85687260     -0.13160712     -1.14301540     -0.04214909 
##      cos_td_rad      sin_td_rad 
##      0.63738395     -0.60406669
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##     -2.48716893      2.85687304     -0.13160658     -1.14301634     -0.04214947 
##      cos_td_rad      sin_td_rad 
##      0.63737835     -0.60406475

## [1] 1640.701
## [1] 0.152156 2.648580
## [1] 16.407
## [1] 17.407

##  subunit          emmean    SE  df asymp.LCL asymp.UCL
##  Judean Highlands -0.454 0.154 Inf    -0.756    -0.153
##  Carmel           -0.586 0.157 Inf    -0.895    -0.277
##  Galilee          -1.597 0.198 Inf    -1.985    -1.210
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## Confidence level used: 0.95
## [1] 174.8348
## [1] 14.11083
## [1] 213.6163
##  contrast                   estimate    SE  df z.ratio p.value
##  Judean Highlands - Carmel     0.132 0.190 Inf   0.692  0.4887
##  Judean Highlands - Galilee    1.143 0.220 Inf   5.195  <.0001
##  Carmel - Galilee              1.011 0.222 Inf   4.560  <.0001
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## P value adjustment: fdr method for 3 tests

Streptopelia decaocto - תור צווארון

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##    -1.436931728     0.157512825     0.290118464    -0.165355750    -0.005583385 
##      cos_td_rad      sin_td_rad 
##     2.033709987    -1.869831164
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##     -1.43693148      0.15751286      0.29011842     -0.16535581     -0.00558339 
##      cos_td_rad      sin_td_rad 
##      2.03370977     -1.86983108

##  subunit          emmean     SE  df asymp.LCL asymp.UCL
##  Judean Highlands   1.15 0.0887 Inf     0.981      1.33
##  Carmel             1.45 0.0852 Inf     1.278      1.61
##  Galilee            0.99 0.0918 Inf     0.810      1.17
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## Confidence level used: 0.95
## [1] 58.4074
## [1] 34.98588
## [1] 17.35109
##  contrast                   estimate    SE  df z.ratio p.value
##  Judean Highlands - Carmel    -0.290 0.123 Inf  -2.359  0.0275
##  Judean Highlands - Galilee    0.165 0.127 Inf   1.297  0.1945
##  Carmel - Galilee              0.455 0.125 Inf   3.637  0.0008
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## P value adjustment: fdr method for 3 tests

Streptopelia turtur - תור מצוי

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##      -4.0568827      -0.3138816       0.8126151       0.2758993      -0.1499697 
##      cos_td_rad      sin_td_rad 
##       2.8254657      -1.6813797
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##      -4.0568824      -0.3138816       0.8126152       0.2758993      -0.1499697 
##      cos_td_rad      sin_td_rad 
##       2.8254655      -1.6813795

## [1] 74.0689
##   [1] 0.34455932 0.33989362 0.33529109 0.33075088 0.32627215 0.32185407
##   [7] 0.31749581 0.31319658 0.30895555 0.30477196 0.30064501 0.29657395
##  [13] 0.29255802 0.28859646 0.28468855 0.28083356 0.27703077 0.27327947
##  [19] 0.26957896 0.26592857 0.26232761 0.25877541 0.25527131 0.25181465
##  [25] 0.24840481 0.24504114 0.24172301 0.23844982 0.23522095 0.23203580
##  [31] 0.22889378 0.22579431 0.22273681 0.21972071 0.21674545 0.21381048
##  [37] 0.21091526 0.20805923 0.20524188 0.20246269 0.19972112 0.19701668
##  [43] 0.19434886 0.19171716 0.18912110 0.18656020 0.18403397 0.18154195
##  [49] 0.17908367 0.17665868 0.17426653 0.17190677 0.16957897 0.16728268
##  [55] 0.16501749 0.16278298 0.16057872 0.15840431 0.15625934 0.15414342
##  [61] 0.15205615 0.14999714 0.14796602 0.14596240 0.14398591 0.14203618
##  [67] 0.14011286 0.13821557 0.13634398 0.13449774 0.13267649 0.13087991
##  [73] 0.12910765 0.12735939 0.12563481 0.12393358 0.12225538 0.12059991
##  [79] 0.11896685 0.11735591 0.11576679 0.11419918 0.11265280 0.11112736
##  [85] 0.10962257 0.10813816 0.10667385 0.10522937 0.10380445 0.10239883
##  [91] 0.10101223 0.09964442 0.09829512 0.09696410 0.09565110 0.09435588
##  [97] 0.09307820 0.09181782 0.09057451 0.08934803
## [1] 0.740689
## [1] 3.856373

Troglodytes troglodytes - גדרון מובהק

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##     -3.84236334     -1.19509372     -0.04556621      0.45798127     -0.10261925 
##      cos_td_rad      sin_td_rad 
##      2.44609621     -3.55529291
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##     -3.84238142     -1.19509312     -0.04556577      0.45798207     -0.10261927 
##      cos_td_rad      sin_td_rad 
##      2.44610919     -3.55530696

## [1] 0.5981707 0.1810518
## [1] 230.3865
## [1] 0.5981707 0.1810518
## [1] 2.303865
## [1] 3.303865

Garrulus glandarius - עורבני

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##     -1.11756667      0.25786121      0.33158628     -0.69131910     -0.05805483 
##      cos_td_rad      sin_td_rad 
##      1.00690364     -0.79489729
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##     -1.43693148      0.15751286      0.29011842     -0.16535581     -0.00558339 
##      cos_td_rad      sin_td_rad 
##      2.03370977     -1.86983108

##  subunit           emmean    SE  df asymp.LCL asymp.UCL
##  Judean Highlands -0.0865 0.135 Inf  -0.35092     0.178
##  Carmel            0.2451 0.126 Inf  -0.00262     0.493
##  Galilee          -0.7778 0.161 Inf  -1.09430    -0.461
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## Confidence level used: 0.95
## [1] 178.1249
## [1] 39.31955
## [1] 99.6309
##  contrast                   estimate    SE  df z.ratio p.value
##  Judean Highlands - Carmel    -0.332 0.185 Inf  -1.796  0.0725
##  Judean Highlands - Galilee    0.691 0.210 Inf   3.294  0.0015
##  Carmel - Galilee              1.023 0.205 Inf   4.993  <.0001
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## P value adjustment: fdr method for 3 tests

Parus major - ירגזי מצוי

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##     -1.94131820      0.21506433      1.02233232      0.59434990     -0.06096473 
##      cos_td_rad      sin_td_rad 
##      0.90791148     -2.05889505
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##     -1.94131734      0.21506408      1.02233220      0.59435010     -0.06096475 
##      cos_td_rad      sin_td_rad 
##      0.90791116     -2.05889431

##  subunit          emmean    SE  df asymp.LCL asymp.UCL
##  Judean Highlands -0.275 0.145 Inf   -0.5580   0.00871
##  Carmel            0.748 0.123 Inf    0.5071   0.98826
##  Galilee           0.320 0.132 Inf    0.0618   0.57767
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## Confidence level used: 0.95
## [1] 53.41861
## [1] 178.1527
## [1] 81.30309
##  contrast                   estimate    SE  df z.ratio p.value
##  Judean Highlands - Carmel    -1.022 0.189 Inf  -5.397  <.0001
##  Judean Highlands - Galilee   -0.594 0.195 Inf  -3.043  0.0035
##  Carmel - Galilee              0.428 0.180 Inf   2.380  0.0173
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## P value adjustment: fdr method for 3 tests

Corvus.monedula - קאק

##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##      0.82864990     -0.01170426     -2.01179194     -3.12892756     -0.02734277 
##      cos_td_rad      sin_td_rad 
##     -0.01871584      0.59352267
##     (Intercept) settlementsNear   subunitCarmel  subunitGalilee         year_ct 
##      0.82895652     -0.01174288     -2.01172173     -3.12898154     -0.02737127 
##      cos_td_rad      sin_td_rad 
##     -0.01884941      0.59362987

##  subunit          emmean    SE  df asymp.LCL asymp.UCL
##  Judean Highlands   0.33 0.377 Inf    -0.409     1.070
##  Carmel            -1.68 0.416 Inf    -2.497    -0.866
##  Galilee           -2.80 0.506 Inf    -3.790    -1.808
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## Confidence level used: 0.95
## [1] 206.4854
## [1] 646.3317
## [1] 2187.398
##  contrast                   estimate    SE  df z.ratio p.value
##  Judean Highlands - Carmel      2.01 0.562 Inf   3.581  0.0005
##  Judean Highlands - Galilee     3.13 0.630 Inf   4.963  <.0001
##  Carmel - Galilee               1.12 0.654 Inf   1.707  0.0878
## 
## Results are averaged over the levels of: settlements 
## Results are given on the log (not the response) scale. 
## P value adjustment: fdr method for 3 tests