Introduction

Juvenile fish are important prey for marine birds and mammals. During June 2021, three juvenile fish were caught during surface water sampling near the Isles of Shoals and were identified as red hake (Urophycis chuss), white hake (Urophycis tenuis) or fourbeard rockling (Enchelyopus cimbrius), which are morphologically similar species (Marancik et al. 2020). These three species are prey of the terns (Gemma Clucas, Elizabeth Craig, pers. comm.) and red and white hake are prey of the seals (Veo et al., 2021) found at the Isles of Shoals. Juvenile red and white hake spend their first months drifting at or near surface water, sometimes under floating eelgrass or rockweed, reaching ~10 cm by fall (Steiner et al., 1982). Juvenile fourbeard rockling also spend their first months at or near surface water (Bigelow and Schroeder, 1953). Red and white hake may be less nutritious than other seal and tern prey such as Atlantic herring (Clupea harengus) and American sand lance (Ammodytes americanus), yet if they are common at the water surface, may make an important contribution to these predators’ diets, similar to prey patterns for Steller sea lions in southeast Alaska (Sigler et al., 2009). During the 2021 sampling, eight neuston net tows were completed during surface water sampling. The intent of this project is to continue this sampling and to characterize the juvenile fish near the Isles of Shoals.

Objective

Characterize the juvenile fish near the Isles of Shoals.

Methods

A neuston net was deployed to sample juvenile fish at the water surface. The net is a standard configuration of 1-m wide, 0.5-m deep and 3-m long with 330-micron mesh and cod end. This neuston net configuration is the standard net used by the NOAA Northeast Fisheries Science Center (pers. comm. Jerry Prezioso, NOAA Northeast Fisheries Science Center, 18 March 2020). Although neuston nets are not routinely used during standard surveys of the northeast U.S. shelf, using this net provides comparable data to data collected by the NOAA Northeast Fisheries Science Center. The neuston net was deployed from a davit on the R/V John B. Heiser and occasionally, from the vessel crane on the R/V John M. Kingsbury. The net was deployed off the side of the vessel, 5-minute tow duration, with the vessel turning slowly during the tow to keep the net outside of the vessel wake. Following retrieval, a washdown pump and hose were used to the wash net contents into the cod end and to clean the net. Following collection, the samples were stored in 1-liter wide mouth plastic jars, returned to the lab and if necessary, refrigerated overnight before processing the samples the next day.

Sampling occurred during low light periods (near dusk) when juvenile fish are more likely to be near the surface. In 2022, sampling primarily occurred in a restricted passage where fish may be concentrated (the channel between Smuttynose and Appledore islands, where juvenile fish were found in 2021). In 2023, sampling primary occurred near the tern colony at White and Seavey islands and near the seal colony at Duck Island and adjacent ledges. A sampling period consisted of three tows, lasted one hour, and occurred weekly for eight weeks in 2022 and for nine weeks in 2023.

In the lab, samples were filtered through a 4-mm sieve to remove debris (e.g., algae) and into a 330-micron sieve (the same mesh size as the neuston net). Fish were individually picked from the 330-micron sieve and transferred to a watch glass for identification. A dissecting scope was used to identify species and to count and length the fish. Images of the catch were recorded often in 2022 and routinely in 2023 for species verification and electronic length measurements using the open-source software Image J. Zooplankton usually were identified to general taxa (e.g., copepods) and their abundance approximated (e.g., many).

In addition, a beach seine was deployed to sample another habitat where juvenile fish commonly are found, coastal eelgrass habitats. The beach seine is a standard configuration with a 37-m long variable-mesh beach seine that tapered from 5-m wide at the center to 1-m wide at the ends. Outer panels were each 10-m of 32-mm stretched mesh, intermediate panels were each 4-m of 6-mm square mesh, and the bunt was 9-m of 3.2-mm square mesh. The seine had a lead line and a float line so that the bottom contacted the substrate and the top floated. This beach seine is a standard configuration used on the U.S. and Canada Pacific coast and is an effective tool for sampling eelgrass habitat (e.g., Johnson et al. 2003). This beach seine is somewhat larger than beach seines used by NH Fish and Game Department to survey estuarine juvenile finfish (30.5 m long and 1.8 m high with 6.4 mm mesh, with a bag where the height, width, and depth each measure 1.8 m) (NH Fish and Game, 2022).

We set the beach seine as a ”round haul”. The net was deployed off the bow of an inflatable. Seine deployment began at the beach, setting a line attached to one end of the seine, motoring away from the beach for 30-40 m, turning parallel to the beach and starting the seine deployment. Once the seine was deployed (parallel to the beach), a line attached to the other end of the seine was brought back to the beach. At this point, the seine lies parallel to the beach and 30-40 m away. Three to four people began hauling each line onto the beach (six to eight people total), steadily pulling the seine through the water until the entire seine is on the lower beach. Following retrieval, the catch was removed from the seine and placed in buckets. The catch was identified to species, counted and for a subsample of each fish species, the length was measured. Beach seine sampling occurred during daylight hours following a low tide, so that the seine could reach and sample the eel grass habitat. Two locations were sampled, Fort Foster Beach, Maine and New Castle Beach, New Hampshire in 2022 and New Castle beach in 2023. A sampling period typically consisted of one beach seine deployment.

Results and Discussion

Species identification

Juvenile fish identification can be difficult, especially for the earliest larval stages. Physical characteristics change quickly during early life and species often are morphologically similar (e.g., Marancik et al. 2020). For example, larvae of red and white hake Urophycis sp. have 2-3 pelvic fin rays whereas fourbeard rockling have 4-7 (Fahay 1983), seemingly diagnostic, but sometimes the number of rays can be difficult to count if fin membranes are torn (pers. observation). Some species were initially misidentified in 2022 and species identification improved in 2023. Taxonomic experts at NOAA Northeast Fisheries Science Center (Katey Marancik and Elizabeth Broughton) have been expert and willing teachers. Once fish advance from the larval to the juvenile fish stage, identification is much easier.

Neuston tows

A total of 31 neuston tows were completed during June and July 2022 at approximately weekly intervals (Figure 1). Neuston tows were completed south (in the channel between Appledore and Smuttynose islands) and east of Appledore Island (Figure 2). The most common fish species caught were red hake or white hake (hereafter “hake”) or fourbeard rockling (Figure 3). Larvae and juveniles of red and white hake are similar (Fahay 1983, Marancik et al. 2020) and were not differentiated in this study. Larvae and juveniles of fourbeard rockling also are similar to hake and were distinguished from one another in 2023 but not in 2022. Several individuals of two other species occasionally were caught, cunner (Tautogolabrus adspersus) in early July and Atlantic mackerel (Scomber scombrus) in late July. The maximum number of fish caught in one sampling day (typically three neuston tows) was 94 (Figure 3). Fish catches were minimal until the fourth week of June and remained elevated thereafter. The length range of all species was 5 - 28 mm TL; most fish were 5-20 mm TL (Figure 4).

A total of 28 neuston tows were completed during June, July and August 2023 at approximately weekly intervals (Figure 1). Neuston tows were completed near the tern colony at White and Seavey islands and near the seal colony at Duck Island and adjacent ledges (Figure 2). The most common fish species caught were hake or fourbeard rockling (Figure 3). The maximum number of fish caught in one sampling day (three neuston tows) was 11. Fish catches were initially nil, increased in mid-July and remained elevated thereafter. Fish catches were smaller in 2023 than in 2022. The length range of all species was 2 - 36 mm TL; most fish were 8-21 mm TL (Figure 4).

Beach seines

A total of five beach seine sets were completed during 2021-2023, two each in 2021 and 2023 and one in 2022 (Figure 1). The beach seine sets were located at the mouth of the Piscataqua River. Most of the beach seines were completed at New Castle Beach, New Hampshire and one, in 2021, was completed at Fort Foster Beach, Maine (Figure 2). The most common fish species caught was sand lance, due to one large sample of about 2,000 juveniles (Figure 3). Other common fish species were Atlantic herring, Atlantic tomcod (Microgadus tomcod) and rainbow smelt (Osmerus mordax). The maximum number of fish caught in one beach seine set was about 2,000 (Figure 2). Only one juvenile flatfish was caught at Fort Foster Beach in 2021 whereas on the same day, about 20 juvenile fish were caught at New Castle Beach. Too few beach seine sets were completed to measure seasonal catch patterns (Figure 1). The length range of all species was 15 - 210 mm TL; most fish were 30-120 mm TL (Figure 4).

Figures 1-4

Figure 1 Sampling effort

Sampling effort (number of gear deployments) by gear type (neuston net near the Isles of Shoals; beach seine in the mouth of the Piscataqua River), day of year and year during the summers of 2021-2023.

Figure 2 Sampling locations

Sample locations by gear type (neuston net primarily near the Isles of Shoals, upper panel; beach seine in the mouth of the Piscataqua River, lower panel) during the summers of 2021-2023.

Figure 3 Fish catches

Fish catch by species, day of year and year caught by gear type (neuston net near the Isles of Shoals, upper panel; beach seine in the mouth of the Piscataqua River, lower panel) during the summers of 2021-2023.

Figure 4 Fish lengths

Fish total length (mm) by species, day of year and year caught by gear type (neuston net near the Isles of Shoals, upper panel; beach seine in the mouth of the Piscataqua River, lower panel) during the summers of 2021-2023.

Habitat comparison

Two gear types were used to sample two different habitats. The neuston net was used to sample surface water near the Isles of Shoals. The beach seine was used to sample coastal eelgrass habitat in the mouth of the Piscataqua River. The species and sizes caught by the two sampling gears overlapped little. Only one species, cunner, was caught on at least two occasions by both neuston net and beach seine (Figure 5). Two species, stickleback (Gasterosteus aculeatus) and northern pipefish (Syngnathus fuscus) were sometimes caught by beach seine but only once by neuston net. In addition, fish caught by neuston net were smaller than fish caught by beach seine (Figure 6). These differences reflect the different species and life stages inhabiting the habitats sampled by the two gears:
* larvae and early young of the year juveniles in the surface water sampled by the neuston net;
* young of the year and likely age one and possibly older juveniles in the eelgrass habitat sampled by the beach seine.

Figures 5-6

Figure 5 Habitat comparison fish catches

Fish catch by species and gear type (beach seine samples of coastal eelgrass habitat in the mouth of the Piscataqua River, left panel; neuston net samples of surface water near the Isles of Shoals, right panel) during the summers of 2021-2023.

Figure 6 Habitat comparison fish lengths

Fish total length (mm) by gear type (beach seine samples of coastal eelgrass habitat in the mouth of the Piscataqua River, upper panel; neuston net samples of surface water near the Isles of Shoals, lower panel) during the summers of 2021-2023.

Collaboration

The juvenile fish project complements existing research projects at the Isles of Shoals and nearby and provided opportunities for collaboration. These include Aliya Caldwell’s Ph.D. research on tern foraging, long-term tern research at White and Seavey islands led by Dr. Elizabeth Craig, long-term seal research at Duck Island led by Dr. Andrea Bogomolni and the fisheries internship collecting underwater camera information on fish abundance and diversity, which has been led by Dr. Easton White, Dr. Jennifer Dijkstra, Owen Nichols, and Dr. Lindsey Williams.

Product

Observations of the abundance and size of juvenile fish by week and location. See the appendix for the database of sample position, catch, and length.

Acknowledgements

Thanks to the Shoals Marine Laboratory for funding this research and providing me with the opportunity to be a Scientist in Residence for 2022 and 2023. The undergraduate researchers were great and I had fun watching them grow as scientists and guiding their research. Thanks to the students, undergraduate researchers, staff and faculty who helped to catch and process samples including from the Marine Ecosystem Research and Management class which I teach with Drs. Ebett Siddon and Chris Siddon. Thanks also to the taxonomic experts at the NOAA Northeast Fisheries Science Center, Katey Marancik and Elizabeth Broughton, who have been expert and willing teachers, and an oceanographer at the NOAA Northeast Fisheries Science Center, Jerry Prezioso, who ably answered questions about the NOAA Northeast Fisheries Science Center sampling methods, data and data availability. Thanks also to Dr. Rebecca Atkins, my fellow Scientist in Residence in 2023, for introducing me to the interactive plotting used in the html version of this report.

References

Bigelow, H.B. and Schroeder, W.C., 1953. Fishes of the Gulf of Maine (No. 592). US Government Printing Office. link

Fahay, M.P. 1983. Guide to the early stages of marine fishes occurring in the western North Atlantic Ocean, Cape Hatteras to the southern Scotian Shelf. J. Northwest Atlantic Fishery Science, 4:1-423. Updated online version available at link

Marancik, K.E., Richardson, D.E. and Konieczna, M., 2020. Updated morphological descriptions of the larval stage of Urophycis (Family: Phycidae) from the Northeast United States continental shelf. Copeia, 108(1), pp.83-90. link

New Hampshire Fish and Game Department. 2022. Estuarine survey of juvenile finfish. link

Johnson, S.W., Murphy, M.L., Csepp, D.J., Harris, P.M. and Thedinga, J.F. 2003. A survey of fish assemblages in eelgrass and kelp habitats of southeastern Alaska. US Dept Commerce, NOAA Tech. Memo. NMFS-AFSC-139, 39 p. link

Sigler, M.F., Tollit, D.J., Vollenweider, J.J., Thedinga, J.F., Csepp, D.J., Womble, J.N., Wong, M.A., Rehberg, M.J. and Trites, A.W. 2009. Steller sea lion foraging response to seasonal changes in prey availability. Mar. Ecol. Prog. Ser. 388: 243-261. link

Steiner, W.W., Luczkovich, J.J. and Olla, B.L. 1982. Activity, Shelter Usage, Growth and Recruitment of Juvenile Red Hake Urophycis chuss. Mar. Ecol. Prog. Ser. 7: 125-135. link

Veo, J., Berube, J., Bogomolni, A., Lysiak, N. and Sette, L. 2022. Characterizing the diet of stranded harbor seals (Phoca vitulina concolor) in the Gulf of Maine. Presentation to the Shoals Marine Laboratory class “Marine Ecosystem Research and Management”.

Appendix

Sample locations

Sample locations by gear type (neuston net near the Isles of Shoals; beach seine in the mouth of the Piscataqua River) during the summers of 2021-2023.

Event Gear Date Start time (HHMM) Latitude (DD.DDDDD) Longitude (DD.DDDDD) Bottom depth (m)
20 neuston net 2021-06-17 1345 42.98733 -70.60583 36
21 neuston net 2021-06-17 1404 42.98667 -70.60183 30
22 neuston net 2021-06-17 1423 42.98690 -70.60017 30
25 neuston net 2021-06-19 1940 42.98715 -70.60242 40
26 neuston net 2021-06-19 2005 42.98430 -70.60653 12
27 neuston net 2021-06-19 2018 42.98267 -70.61588 14
30 neuston net 2021-06-24 1143 43.06800 -70.70985 10
31 neuston net 2021-06-24 1156 43.06883 -70.71063 8
35 neuston net 2022-06-04 1356 42.98597 -70.60865 12
36 neuston net 2022-06-04 1419 42.98558 -70.61025 6
37 neuston net 2022-06-04 1431 42.98348 -70.61420 14
38 neuston net 2022-06-04 1445 42.98257 -70.61368 7
39 neuston net 2022-06-04 1519 42.99150 -70.61018 14
40 neuston net 2022-06-04 1529 42.99397 -70.60957 28
41 neuston net 2022-06-04 1542 42.99242 -70.60562 27
42 neuston net 2022-06-07 845 42.98527 -70.60958 10
43 neuston net 2022-06-07 901 42.98380 -70.61270 12
44 neuston net 2022-06-11 1857 42.98432 -70.61303 8
45 neuston net 2022-06-11 1908 42.98385 -70.61460 13
46 neuston net 2022-06-11 1920 42.98720 -70.60847 13
47 neuston net 2022-06-11 1930 42.98407 -70.60977 15
48 neuston net 2022-06-16 1926 42.98360 -70.61130 10
49 neuston net 2022-06-16 1941 42.98392 -70.61137 14
50 neuston net 2022-06-16 1954 42.98362 -70.61190 12
51 neuston net 2022-06-21 1858 42.98278 -70.61408 12
52 neuston net 2022-06-21 1908 42.98277 -70.61410 21
53 neuston net 2022-06-21 1921 42.98565 -70.60933 12
55 neuston net 2022-07-02 1859 42.98265 -70.61888 18
56 neuston net 2022-07-02 1913 42.98360 -70.61020 8
57 neuston net 2022-07-02 1923 42.98733 -70.60747 13
58 neuston net 2022-07-11 1912 42.98345 -70.61237 8
59 neuston net 2022-07-11 1930 42.98530 -70.60178 24
60 neuston net 2022-07-11 1938 42.98417 -70.60613 7
63 neuston net 2022-07-16 1917 42.98383 -70.61117 13
64 neuston net 2022-07-16 1928 42.98467 -70.60767 21
65 neuston net 2022-07-16 1943 42.98333 -70.61500 12
66 neuston net 2022-07-23 1903 42.98217 -70.62033 NA
67 neuston net 2022-07-23 1915 42.98360 -70.61200 NA
68 neuston net 2022-07-23 1926 42.98603 -70.60777 NA
69 neuston net 2023-06-11 1905 42.99198 -70.61875 17
70 neuston net 2023-06-11 1917 42.99617 -70.63092 17
71 neuston net 2023-06-11 1927 42.99622 -70.62693 31
72 neuston net 2023-06-17 1905 42.98415 -70.61935 15
73 neuston net 2023-06-17 1912 42.98162 -70.61913 26
74 neuston net 2023-06-17 1925 42.98018 -70.61648 16
75 neuston net 2023-06-23 1927 42.96913 -70.62217 11
76 neuston net 2023-06-23 1936 42.96945 -70.62210 12
77 neuston net 2023-06-23 1947 42.96945 -70.62210 11
79 neuston net 2023-07-01 1921 42.99975 -70.59697 26
80 neuston net 2023-07-01 1934 43.00265 -70.59855 18
81 neuston net 2023-07-01 1947 43.00705 -70.59780 24
83 neuston net 2023-07-08 1930 42.96892 -70.62050 21
84 neuston net 2023-07-08 1942 42.97050 -70.62508 14
85 neuston net 2023-07-08 1953 42.96912 -70.62880 12
88 neuston net 2023-07-15 1930 43.00700 -70.60575 12
89 neuston net 2023-07-15 1940 43.00705 -70.59940 20
90 neuston net 2023-07-15 1952 43.00443 -70.59665 15
93 neuston net 2023-07-26 1930 42.96498 -70.62620 14
94 neuston net 2023-07-26 1946 42.96793 -70.63043 23
95 neuston net 2023-07-26 1956 42.96928 -70.62740 7
100 neuston net 2023-07-29 1900 43.00213 -70.60038 7
101 neuston net 2023-07-29 1911 43.00822 -70.60098 9
102 neuston net 2023-07-29 1927 43.00520 -70.60998 14
106 neuston net 2023-08-05 1938 42.96962 -70.62200 13
107 neuston net 2023-08-05 1948 42.97073 -70.62442 18
108 neuston net 2023-08-05 1959 42.96740 -70.63102 9
110 neuston net 2023-08-12 2130 42.99249 -70.61990 21
33 beach seine 2021-06-24 1315 43.06738 -70.69775 3
34 beach seine 2021-06-24 1425 43.06738 -70.69775 2
54 beach seine 2022-06-23 1300 43.06795 -70.71301 3
82 beach seine 2023-07-07 1030 43.06795 -70.71301 3
105 beach seine 2023-08-05 930 43.06795 -70.71301 3

Fish catches

Fish catches by gear type (neuston net near the Isles of Shoals; beach seine in the mouth of the Piscataqua River) during the summers of 2021-2023.

Event Gear Date Species Count
20 neuston net 2021-06-17 fish unident 1
25 neuston net 2021-06-19 hake/rockling 2
26 neuston net 2021-06-19 hake/rockling 1
33 beach seine 2021-06-24 rainbow smelt 2
33 beach seine 2021-06-24 sandlance 16
33 beach seine 2021-06-24 herring 4
43 neuston net 2022-06-07 red/white hake 1
44 neuston net 2022-06-11 hake/rockling 1
49 neuston net 2022-06-16 hake/rockling 1
50 neuston net 2022-06-16 hake/rockling 1
51 neuston net 2022-06-21 hake/rockling 18
51 neuston net 2022-06-21 three-spined stickleback 1
52 neuston net 2022-06-21 hake/rockling 15
53 neuston net 2022-06-21 hake/rockling 3
54 beach seine 2022-06-23 rainbow smelt 2
54 beach seine 2022-06-23 sandlance 2000
54 beach seine 2022-06-23 herring 7
54 beach seine 2022-06-23 red/white hake 1
54 beach seine 2022-06-23 emerald shiner 1
54 beach seine 2022-06-23 lumpfish 1
54 beach seine 2022-06-23 northern pipefish 1
54 beach seine 2022-06-23 tomcod 4
54 beach seine 2022-06-23 stickleback 1
54 beach seine 2022-06-23 pollock 1
55 neuston net 2022-07-02 hake/rockling 23
55 neuston net 2022-07-02 cunner 2
55 neuston net 2022-07-02 daubed shanny 1
56 neuston net 2022-07-02 hake/rockling 12
57 neuston net 2022-07-02 hake/rockling 22
58 neuston net 2022-07-11 fish unident 16
59 neuston net 2022-07-11 hake/rockling 32
59 neuston net 2022-07-11 cunner 36
59 neuston net 2022-07-11 mackerel 6
60 neuston net 2022-07-11 fish unident 4
65 neuston net 2022-07-16 fish unident 30
66 neuston net 2022-07-23 hake/rockling 6
66 neuston net 2022-07-23 mackerel 14
67 neuston net 2022-07-23 hake/rockling 15
67 neuston net 2022-07-23 cunner 1
68 neuston net 2022-07-23 hake/rockling 8
68 neuston net 2022-07-23 mackerel 2
80 neuston net 2023-07-01 red/white hake 3
81 neuston net 2023-07-01 red/white hake 1
82 beach seine 2023-07-07 rainbow smelt 3
82 beach seine 2023-07-07 sandlance 23
82 beach seine 2023-07-07 herring 60
82 beach seine 2023-07-07 three-spined stickleback 13
82 beach seine 2023-07-07 northern pipefish 2
82 beach seine 2023-07-07 tomcod 65
82 beach seine 2023-07-07 pollock 7
82 beach seine 2023-07-07 cunner 1
82 beach seine 2023-07-07 winter flounder 1
89 neuston net 2023-07-15 fourbeard rockling 1
90 neuston net 2023-07-15 fourbeard rockling 5
95 neuston net 2023-07-26 fourbeard rockling 1
100 neuston net 2023-07-29 hake/rockling 3
100 neuston net 2023-07-29 fourbeard rockling 2
101 neuston net 2023-07-29 hake/rockling 3
101 neuston net 2023-07-29 northern pipefish 1
101 neuston net 2023-07-29 fourbeard rockling 2
105 beach seine 2023-08-05 rainbow smelt 17
105 beach seine 2023-08-05 northern pipefish 11
105 beach seine 2023-08-05 tomcod 22
105 beach seine 2023-08-05 stickleback 16
105 beach seine 2023-08-05 pollock 4
105 beach seine 2023-08-05 cunner 10
105 beach seine 2023-08-05 sculpin 4
106 neuston net 2023-08-05 fourbeard rockling 2
107 neuston net 2023-08-05 fourbeard rockling 2
108 neuston net 2023-08-05 fourbeard rockling 1
110 neuston net 2023-08-12 red/white hake 6
110 neuston net 2023-08-12 fourbeard rockling 2

Fish lengths

Fish total lengths (mm) by gear type (neuston net near the Isles of Shoals; beach seine in the mouth of the Piscataqua River) during the summers of 2021-2023.

Event Gear Species Length (mm) Count
25 neuston net hake/rockling 32 1
25 neuston net hake/rockling 8 1
26 neuston net hake/rockling 35 1
33 beach seine herring 40 1
33 beach seine herring 40 1
33 beach seine herring 40 1
33 beach seine herring 40 1
33 beach seine herring 42 1
33 beach seine sandlance 45 1
33 beach seine sandlance 50 1
33 beach seine sandlance 50 1
33 beach seine sandlance 52 1
33 beach seine sandlance 60 1
33 beach seine sandlance 60 1
33 beach seine sandlance 60 1
33 beach seine sandlance 60 1
33 beach seine sandlance 60 1
33 beach seine sandlance 68 1
33 beach seine sandlance 68 1
33 beach seine sandlance 70 1
33 beach seine sandlance 70 1
33 beach seine sandlance 70 1
33 beach seine sandlance 70 1
33 beach seine sandlance 70 1
33 beach seine rainbow smelt 30 1
33 beach seine rainbow smelt 30 1
43 neuston net red/white hake 8 1
44 neuston net hake/rockling 10 1
49 neuston net hake/rockling 10 1
50 neuston net hake/rockling 6 1
51 neuston net three-spined stickleback 28 1
51 neuston net hake/rockling 6 1
51 neuston net hake/rockling 6 1
51 neuston net hake/rockling 6 1
51 neuston net hake/rockling 6 1
51 neuston net hake/rockling 13 1
51 neuston net hake/rockling 13 1
51 neuston net hake/rockling 13 1
51 neuston net hake/rockling 14 1
51 neuston net hake/rockling 14 1
51 neuston net hake/rockling 14 1
51 neuston net hake/rockling 15 1
51 neuston net hake/rockling 15 1
51 neuston net hake/rockling 15 1
51 neuston net hake/rockling 16 1
51 neuston net hake/rockling 17 1
51 neuston net hake/rockling 17 1
51 neuston net hake/rockling 18 1
51 neuston net hake/rockling 20 1
52 neuston net hake/rockling 10 1
52 neuston net hake/rockling 11 1
52 neuston net hake/rockling 11 1
52 neuston net hake/rockling 11 1
52 neuston net hake/rockling 12 1
52 neuston net hake/rockling 12 1
52 neuston net hake/rockling 13 1
52 neuston net hake/rockling 13 1
52 neuston net hake/rockling 14 1
52 neuston net hake/rockling 14 1
52 neuston net hake/rockling 15 1
52 neuston net hake/rockling 15 1
52 neuston net hake/rockling 16 1
52 neuston net hake/rockling 16 1
52 neuston net hake/rockling 18 1
52 neuston net hake/rockling 23 1
53 neuston net hake/rockling 11 1
53 neuston net hake/rockling 12 1
53 neuston net hake/rockling 16 1
52 neuston net hake/rockling 28 1
54 beach seine tomcod 210 1
54 beach seine tomcod 95 1
54 beach seine pollock 85 1
54 beach seine herring 80 1
54 beach seine herring 92 1
54 beach seine herring 90 1
54 beach seine herring 110 1
54 beach seine herring 80 1
54 beach seine herring 100 1
54 beach seine emerald shiner 110 1
54 beach seine lumpfish 80 1
54 beach seine northern pipefish 130 1
54 beach seine tomcod 90 1
54 beach seine tomcod 80 1
54 beach seine red/white hake 60 1
54 beach seine herring 97 1
54 beach seine stickleback 40 1
54 beach seine red/white hake 63 1
54 beach seine sandlance 47 1
54 beach seine sandlance 58 1
54 beach seine sandlance 51 1
54 beach seine sandlance 55 1
54 beach seine sandlance 64 1
54 beach seine sandlance 48 1
54 beach seine sandlance 60 1
54 beach seine sandlance 72 1
54 beach seine sandlance 57 1
54 beach seine sandlance 56 1
54 beach seine sandlance 60 1
54 beach seine sandlance 67 1
54 beach seine sandlance 54 1
54 beach seine sandlance 58 1
54 beach seine sandlance 63 1
54 beach seine sandlance 60 1
54 beach seine sandlance 57 1
54 beach seine sandlance 55 1
54 beach seine sandlance 60 1
54 beach seine sandlance 60 1
54 beach seine sandlance 61 1
54 beach seine sandlance 48 1
54 beach seine sandlance 64 1
54 beach seine sandlance 58 1
54 beach seine sandlance 52 1
54 beach seine sandlance 70 1
54 beach seine sandlance 58 1
54 beach seine sandlance 58 1
54 beach seine sandlance 65 1
54 beach seine sandlance 58 1
54 beach seine sandlance 49 1
54 beach seine sandlance 43 1
54 beach seine rainbow smelt 108 1
54 beach seine rainbow smelt 135 1
55 neuston net hake/rockling 11 1
55 neuston net hake/rockling 10 1
55 neuston net hake/rockling 12 1
55 neuston net hake/rockling 11 1
55 neuston net hake/rockling 7 1
55 neuston net hake/rockling 5 1
55 neuston net hake/rockling 7 1
55 neuston net hake/rockling 8 1
55 neuston net hake/rockling 8 1
55 neuston net hake/rockling 7 1
55 neuston net hake/rockling 8 1
55 neuston net hake/rockling 8 1
55 neuston net hake/rockling 9 1
55 neuston net hake/rockling 8 1
55 neuston net hake/rockling 6 1
55 neuston net hake/rockling 6 1
55 neuston net hake/rockling 7 1
55 neuston net hake/rockling 9 1
55 neuston net hake/rockling 7 1
55 neuston net hake/rockling 6 1
55 neuston net hake/rockling 6 1
55 neuston net cunner 8 1
55 neuston net cunner 8 1
55 neuston net daubed shanny 11 1
56 neuston net hake/rockling 7 1
56 neuston net hake/rockling 8 1
56 neuston net hake/rockling 8 1
56 neuston net hake/rockling 9 1
56 neuston net hake/rockling 9 1
56 neuston net hake/rockling 10 1
56 neuston net hake/rockling 8 1
56 neuston net hake/rockling 8 1
56 neuston net hake/rockling 8 1
56 neuston net hake/rockling 10 1
56 neuston net hake/rockling 10 1
56 neuston net hake/rockling 7 1
56 neuston net hake/rockling 8 1
57 neuston net hake/rockling 8 1
57 neuston net hake/rockling 6 1
57 neuston net hake/rockling 7 1
57 neuston net hake/rockling 5 1
57 neuston net hake/rockling 9 1
57 neuston net hake/rockling 8 1
57 neuston net hake/rockling 7 1
57 neuston net hake/rockling 7 1
57 neuston net hake/rockling 8 1
57 neuston net hake/rockling 8 1
57 neuston net hake/rockling 10 1
57 neuston net hake/rockling 7 1
57 neuston net hake/rockling 8 1
57 neuston net hake/rockling 7 1
57 neuston net hake/rockling 9 1
57 neuston net hake/rockling 8 1
57 neuston net hake/rockling 8 1
57 neuston net hake/rockling 8 1
57 neuston net hake/rockling 8 1
57 neuston net hake/rockling 10 1
57 neuston net hake/rockling 7 1
58 neuston net fish unident 9 1
58 neuston net fish unident 10 1
58 neuston net fish unident 8 1
58 neuston net fish unident 9 1
58 neuston net fish unident 7 1
58 neuston net fish unident 11 1
58 neuston net fish unident 10 1
58 neuston net fish unident 11 1
58 neuston net fish unident 9 1
58 neuston net fish unident 8 1
58 neuston net fish unident 10 1
58 neuston net fish unident 12 1
58 neuston net fish unident 10 1
58 neuston net fish unident 8 1
58 neuston net fish unident 9 1
58 neuston net fish unident 12 1
58 neuston net fish unident 8 1
60 neuston net fish unident 11 1
60 neuston net fish unident 10 1
60 neuston net fish unident 15 1
60 neuston net fish unident 11 1
59 neuston net cunner 12 1
59 neuston net cunner 12 1
59 neuston net cunner 10 1
59 neuston net cunner 11 1
59 neuston net cunner 8 1
59 neuston net cunner 7 1
59 neuston net cunner 10 1
59 neuston net cunner 12 1
59 neuston net cunner 9 1
59 neuston net cunner 11 1
59 neuston net cunner 14 1
59 neuston net cunner 10 1
59 neuston net cunner 15 1
59 neuston net cunner 8 1
59 neuston net cunner 14 1
59 neuston net cunner 10 1
59 neuston net cunner 14 1
59 neuston net cunner 13 1
59 neuston net hake/rockling 11 1
59 neuston net hake/rockling 11 1
59 neuston net hake/rockling 10 1
59 neuston net hake/rockling 11 1
59 neuston net hake/rockling 10 1
59 neuston net hake/rockling 12 1
59 neuston net hake/rockling 10 1
59 neuston net hake/rockling 13 1
59 neuston net hake/rockling 12 1
59 neuston net hake/rockling 19 1
59 neuston net hake/rockling 11 1
59 neuston net hake/rockling 10 1
59 neuston net hake/rockling 13 1
59 neuston net hake/rockling 12 1
59 neuston net mackerel 8 1
59 neuston net mackerel 8 1
65 neuston net fish unident 8 1
65 neuston net fish unident 15 1
65 neuston net fish unident 11 1
65 neuston net fish unident 12 1
65 neuston net fish unident 12 1
65 neuston net fish unident 12 1
65 neuston net fish unident 11 1
65 neuston net fish unident 12 1
65 neuston net fish unident 10 1
65 neuston net fish unident 8 1
65 neuston net fish unident 11 1
65 neuston net fish unident 8 1
65 neuston net fish unident 13 1
65 neuston net fish unident 9 1
65 neuston net fish unident 11 1
65 neuston net fish unident 9 1
65 neuston net fish unident 11 1
65 neuston net fish unident 10 1
65 neuston net fish unident 11 1
65 neuston net fish unident 11 1
65 neuston net fish unident 7 1
65 neuston net fish unident 11 1
65 neuston net fish unident 14 1
65 neuston net fish unident 11 1
65 neuston net fish unident 10 1
65 neuston net fish unident 16 1
65 neuston net fish unident 12 1
65 neuston net fish unident 15 1
65 neuston net fish unident 9 1
65 neuston net fish unident 8 1
65 neuston net fish unident 16 1
66 neuston net hake/rockling 14 1
66 neuston net hake/rockling 17 1
66 neuston net hake/rockling 12 1
66 neuston net hake/rockling 10 1
66 neuston net hake/rockling 12 1
66 neuston net hake/rockling 8 1
66 neuston net cunner 10 1
66 neuston net cunner 8 1
66 neuston net cunner 11 1
66 neuston net cunner 9 1
66 neuston net cunner 12 1
66 neuston net cunner 9 1
66 neuston net cunner 10 1
66 neuston net cunner 9 1
66 neuston net cunner 11 1
66 neuston net cunner 10 1
66 neuston net cunner 8 1
66 neuston net cunner 10 1
66 neuston net cunner 11 1
67 neuston net hake/rockling 8 1
67 neuston net hake/rockling 9 1
67 neuston net hake/rockling 9 1
67 neuston net hake/rockling 9 1
67 neuston net hake/rockling 8 1
67 neuston net hake/rockling 8 1
67 neuston net hake/rockling 8 1
67 neuston net hake/rockling 7 1
67 neuston net hake/rockling 8 1
67 neuston net hake/rockling 8 1
67 neuston net hake/rockling 8 1
67 neuston net cunner 12 1
67 neuston net hake/rockling 10 1
67 neuston net hake/rockling 8 1
67 neuston net hake/rockling 9 1
67 neuston net hake/rockling 9 1
68 neuston net mackerel 10 1
68 neuston net hake/rockling 8 1
68 neuston net hake/rockling 8 1
68 neuston net hake/rockling 8 1
68 neuston net hake/rockling 8 1
68 neuston net mackerel 10 1
68 neuston net hake/rockling 6 1
68 neuston net hake/rockling 14 1
68 neuston net hake/rockling 8 1
68 neuston net hake/rockling 8 1
80 neuston net red/white hake 19 1
80 neuston net red/white hake 11 1
80 neuston net red/white hake 11 1
81 neuston net red/white hake 15 1
82 beach seine herring 48 1
82 beach seine herring 55 1
82 beach seine herring 51 1
82 beach seine herring 55 1
82 beach seine herring 110 1
82 beach seine herring 86 1
82 beach seine herring 79 1
82 beach seine herring 70 1
82 beach seine herring 78 1
82 beach seine herring 85 1
82 beach seine herring 75 1
82 beach seine herring 82 1
82 beach seine herring 77 1
82 beach seine herring 75 1
82 beach seine herring 82 1
82 beach seine herring 72 1
82 beach seine herring 77 1
82 beach seine herring 76 1
82 beach seine herring 77 1
82 beach seine herring 73 1
82 beach seine herring 74 1
82 beach seine herring 82 1
82 beach seine herring 85 1
82 beach seine herring 75 1
82 beach seine herring 73 1
82 beach seine herring 75 1
82 beach seine herring 40 1
82 beach seine herring 37 1
82 beach seine herring 40 1
82 beach seine herring 55 1
82 beach seine herring 48 1
82 beach seine herring 39 1
82 beach seine herring 55 1
82 beach seine herring 48 1
82 beach seine herring 39 1
82 beach seine herring 55 1
82 beach seine herring 45 1
82 beach seine herring 47 1
82 beach seine herring 52 1
82 beach seine herring 38 1
82 beach seine herring 50 1
82 beach seine herring 40 1
82 beach seine herring 45 1
82 beach seine herring 47 1
82 beach seine herring 47 1
82 beach seine herring 50 1
82 beach seine herring 45 1
82 beach seine herring 50 1
82 beach seine herring 35 1
82 beach seine herring 60 1
82 beach seine herring 40 1
82 beach seine herring 40 1
82 beach seine herring 40 1
82 beach seine herring 40 1
82 beach seine herring 45 1
82 beach seine herring 80 1
82 beach seine herring 80 1
82 beach seine herring 80 1
82 beach seine herring 70 1
82 beach seine herring 80 1
82 beach seine pollock 90 1
82 beach seine tomcod 55 1
82 beach seine tomcod 75 1
82 beach seine pollock 60 1
82 beach seine pollock 90 1
82 beach seine tomcod 85 1
82 beach seine tomcod 101 1
82 beach seine pollock 100 1
82 beach seine tomcod 80 1
82 beach seine tomcod 95 1
82 beach seine tomcod 100 1
82 beach seine tomcod 100 1
82 beach seine tomcod 130 1
82 beach seine tomcod 85 1
82 beach seine pollock 90 1
82 beach seine tomcod 100 1
82 beach seine tomcod 95 1
82 beach seine tomcod 100 1
82 beach seine tomcod 85 1
82 beach seine tomcod 100 1
82 beach seine tomcod 95 1
82 beach seine tomcod 95 1
82 beach seine tomcod 98 1
82 beach seine tomcod 85 1
82 beach seine tomcod 70 1
82 beach seine tomcod 90 1
82 beach seine tomcod 95 1
82 beach seine tomcod 95 1
82 beach seine tomcod 90 1
82 beach seine tomcod 82 1
82 beach seine tomcod 80 1
82 beach seine tomcod 97 1
82 beach seine tomcod 80 1
82 beach seine tomcod 95 1
82 beach seine tomcod 90 1
82 beach seine tomcod 90 1
82 beach seine tomcod 100 1
82 beach seine pollock 105 1
82 beach seine tomcod 85 1
82 beach seine tomcod 100 1
82 beach seine tomcod 98 1
82 beach seine tomcod 75 1
82 beach seine tomcod 87 1
82 beach seine tomcod 95 1
82 beach seine tomcod 72 1
82 beach seine tomcod 92 1
82 beach seine tomcod 120 1
82 beach seine tomcod 90 1
82 beach seine tomcod 90 1
82 beach seine tomcod 87 1
82 beach seine pollock 105 1
82 beach seine sandlance 75 1
82 beach seine sandlance 90 1
82 beach seine sandlance 52 1
82 beach seine sandlance 73 1
82 beach seine sandlance 75 1
82 beach seine sandlance 76 1
82 beach seine sandlance 77 1
82 beach seine sandlance 80 1
82 beach seine sandlance 75 1
82 beach seine sandlance 80 1
82 beach seine sandlance 76 1
82 beach seine sandlance 80 1
82 beach seine sandlance 72 1
82 beach seine sandlance 85 1
82 beach seine sandlance 87 1
82 beach seine sandlance 80 1
82 beach seine sandlance 84 1
82 beach seine sandlance 90 1
82 beach seine sandlance 80 1
83 neuston net fish larvae 2 1
83 neuston net fish larvae 2 1
83 neuston net fish larvae 2 1
83 neuston net fish larvae 2 1
84 neuston net fish larvae 2 1
89 neuston net fourbeard rockling 14 1
90 neuston net fourbeard rockling 21 1
90 neuston net fourbeard rockling 20 1
90 neuston net fourbeard rockling 16 1
90 neuston net fourbeard rockling 17 1
90 neuston net fourbeard rockling 17 1
95 neuston net fourbeard rockling 15 1
100 neuston net fourbeard rockling 31 1
100 neuston net fourbeard rockling 13 1
100 neuston net hake/rockling 10 1
100 neuston net hake/rockling 9 1
100 neuston net hake/rockling 10 1
101 neuston net fourbeard rockling 23 1
101 neuston net fourbeard rockling 21 1
101 neuston net hake/rockling 9 1
101 neuston net hake/rockling 8 1
101 neuston net hake/rockling 12 1
105 beach seine tomcod 43 1
105 beach seine tomcod 105 1
105 beach seine tomcod 90 1
105 beach seine tomcod 110 1
105 beach seine tomcod 105 1
105 beach seine tomcod 110 1
105 beach seine tomcod 110 1
105 beach seine tomcod 75 1
105 beach seine tomcod 115 1
105 beach seine tomcod 135 1
105 beach seine tomcod 100 1
105 beach seine tomcod 105 1
105 beach seine tomcod 102 1
105 beach seine tomcod 145 1
105 beach seine tomcod 156 1
105 beach seine tomcod 97 1
105 beach seine tomcod 104 1
105 beach seine tomcod 134 1
105 beach seine tomcod 119 1
105 beach seine tomcod 109 1
105 beach seine tomcod 104 1
105 beach seine tomcod 103 1
105 beach seine cunner 30 1
105 beach seine cunner 28 1
105 beach seine cunner 30 1
105 beach seine cunner 25 1
105 beach seine cunner 27 1
105 beach seine cunner 27 1
105 beach seine cunner 27 1
105 beach seine cunner 15 1
105 beach seine cunner 22 1
105 beach seine cunner 21 1
105 beach seine flatfish unident 55 1
105 beach seine flatfish unident 120 1
105 beach seine flatfish unident 41 1
105 beach seine flatfish unident 106 1
105 beach seine northern pipefish 100 1
105 beach seine northern pipefish 170 1
105 beach seine northern pipefish 111 1
105 beach seine northern pipefish 68 1
105 beach seine northern pipefish 73 1
105 beach seine northern pipefish 145 1
105 beach seine northern pipefish 170 1
105 beach seine northern pipefish 155 1
105 beach seine northern pipefish 76 1
105 beach seine northern pipefish 181 1
105 beach seine northern pipefish 114 1
105 beach seine pollock 100 1
105 beach seine pollock 102 1
105 beach seine pollock 98 1
105 beach seine pollock 101 1
105 beach seine rainbow smelt 62 1
105 beach seine rainbow smelt 45 1
105 beach seine rainbow smelt 44 1
105 beach seine rainbow smelt 34 1
105 beach seine rainbow smelt 35 1
105 beach seine rainbow smelt 50 1
105 beach seine rainbow smelt 23 1
105 beach seine rainbow smelt 51 1
105 beach seine rainbow smelt 50 1
105 beach seine rainbow smelt 55 1
105 beach seine rainbow smelt 55 1
105 beach seine rainbow smelt 60 1
105 beach seine rainbow smelt 40 1
105 beach seine rainbow smelt 45 1
105 beach seine rainbow smelt 36 1
105 beach seine rainbow smelt 64 1
105 beach seine rainbow smelt 55 1
105 beach seine sculpin 45 1
105 beach seine sculpin 54 1
105 beach seine sculpin 47 1
105 beach seine sculpin 44 1
105 beach seine stickleback 45 1
105 beach seine stickleback 34 1
105 beach seine stickleback 35 1
105 beach seine stickleback 44 1
105 beach seine stickleback 36 1
105 beach seine stickleback 35 1
105 beach seine stickleback 46 1
105 beach seine stickleback 55 1
105 beach seine stickleback 39 1
105 beach seine stickleback 36 1
105 beach seine stickleback 48 1
105 beach seine stickleback 37 1
105 beach seine stickleback 49 1
105 beach seine stickleback 45 1
105 beach seine stickleback 35 1
105 beach seine stickleback 46 1
106 neuston net fourbeard rockling 19 1
106 neuston net fourbeard rockling 16 1
107 neuston net fourbeard rockling 15 1
107 neuston net fourbeard rockling 19 1
108 neuston net fourbeard rockling 11 1
110 neuston net fourbeard rockling 36 1
110 neuston net fourbeard rockling 31 1
110 neuston net red/white hake 19 1
110 neuston net red/white hake 21 1
110 neuston net red/white hake 10 1
110 neuston net red/white hake 11 1
110 neuston net red/white hake 15 1
110 neuston net red/white hake 18 1